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Picidae (1 Viewer)

Woodpeckers: H&M-4 (Dickinson and Remsen, 2013) brought attention to the name Chloropicus (Malherbe 1845, type pyrrhogaster), which has priority over Dendropicos (Malherbe 1849, type fuscescens, subspecies lafresnayi).
Well, they did a little bit more than to "bring attention to" this name, in practice. (I should maybe write "unfortunately"...) They designated a type species for the name, which actively made it a senior synonym of Thripias Cabanis & Heine 1863 (and Dendropicos Malherbe 1849 sensu lato), while this name had historically never been treated this way.

Chloropicus Malherbe 1845 [OD]; introduced as a subgenus of Picus.
Incorrect subsequent spelling "Chloropicos" introduced in 1849 by Malherbe himself [here] (presumably to "correct" the "hybrid nature" of the name - χλωρός, "green", Greek + picus, "woodpecker", Latin; "πικος" is not a genuine Greek word either, though, so the name presumably remained a "barbarism" to some). He later reverted to the OS (eg., Malherbe 1862).
Originally included nominal species:
  1. Picus (Chloropicus) Pyrrhogaster Malherbe 1845 [OD] (= Dendropicos (Thripias) pyrrhogaster (Malherbe 1845))
  2. Picus (Chloropicus) Kirkii Malherbe 1845 [OD] (= Veniliornis kirkii (Malherbe 1845))
  3. Picus (Chloropicus) Rufoviridis Malherbe 1845 [OD] (= Campethera maculosa (Valenciennes 1826))
  4. Picus (Chloropicus) Xanthoderus Malherbe 1845 [OD] (= Picus chlorolophus chlorigaster Jerdon 1844)
No original type fixation.

Of these four species, the only one that Malherbe 1862 finally retained in Chloropicus is xanthoderus [here]; pyrrhogaster went to Mesopicus [here]; kirkii too [here]; rufoviridis went to Chrysopicus [here]. The species he included in Chloropicus at this point are species that we now place in Picus or Chrysophlegma.
Gray 1855 [here] and 1870 [here] did not designate a type species, but placed the genus in the synonymy of Gecinus Boie 1830, which he used for the species we now place in Picus. (Views on the type species of Picus have varied quite a bit in the past.)
Stejneger 1886 [here] cited the name from the 1849 subsequent spelling, placed it in the synonymy of Picus Linnaeus 1758, and designated Picus viridis as its type.
Hargitt 1890 [here] cited the name from the 1849 subsequent spelling, placed it in the synonymy of Chrysophlegma Gould 1849, and designated "C. flavinucha" as its type.
Stresemann 1921 [here] cited the name from the original 1845 publication, placed it in the synonymy of Picus Linnaeus 1758, and accepted Stejneger's designation of Picus viridis as its type.
Stuart Baker 1930 [here] cited the name from the original 1845 publication, placed it in the synonymy of Picus Linnaeus 1758, and claimed that Picus viridis was its type by original designation.

Unfortunately, these older type designations are invalid because the designated species was not originally included in the genus. (They were included in 1849, but not in the original 1845 work.) However, the above shows clearly that the name had for a very long time consistently been regarded as a synonym of Picus/Chrysophlegma: this is undoubtedly the reason why Peters 1948 did not use it for a completely other group. And this was not in conflict with the original composition of the genus, as one of the originally included species is nowadays still placed in Picus. This means that, if a type designation was to be done, Picus (Chloropicus) xanthoderus should have been selected, in order to maintain the long-accepted position of the name, and minimize the disruption.

Additionally, I'd be interested if anybody could show me a post-1899 (but pre-2013) work where this name is used as valid. I have not been able to trace any, which suggests that the name should probably have been declared a nomen oblitum, rather than being taken back into use.
 
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OK, help this layman out. First of all, how on Earth did the authors of the paper come about firmly placing mahrattensis in Leiopicus (and stating that the clades are geographically monophyletic), when it certainly seems to be embedded in Dendropicos (or whatever it should be called) or at least jumps around? John Boyd seems to interpret the paper differently, including mahrattensis in "Dendropicos".

Then regarding this new Dendrocopos/Dendropicos/Picoides classification – are there diagnosable morphological differences between the "new" genera? Voice? Anything? And are there any theories than can explain why unrelated geographically overlapping species look so much alike, like in Hairy/Downy, Great Spotted/Middle Spotted, Crimson-breasted/Darjeeling? Is this unique for the woodpeckers?
 
Yellow-crowned Woodpecker

...how on Earth did the authors of the paper come about firmly placing mahrattensis in Leiopicus (and stating that the clades are geographically monophyletic), when it certainly seems to be embedded in Dendropicos (or whatever it should be called) or at least jumps around?
Fuchs & Pons...
... A new taxonomic classification of the pied woodpeckers largely in accordance with our proposition has been recently proposed by Winkler et al. (2014). The main differences with our taxonomic proposition concerns the genus assignment of dorae, auriceps, medius and mahrattensis. We here propose to assign medius, auriceps and dorae to Desertipicus and not to the genera Leoipicus, as suggested by Winkler and Christie (2002b) and Winkler et al. (2014), and mahrattensis to Leiopicus. In our classification, the genus Leiopicus is thus monospecific. The recognition of Desertipicus is justified by the uncertainties concerning the relationships of mahrattensis, the type species of Leiopicus, in our analyses. If further studies confirm the sister-group relationship between mahrattensis and the three Desertipicus species, we would recommend lumping Desertipicus into Leiopicus. ...
 
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And are there any theories than can explain why unrelated geographically overlapping species look so much alike, like in Hairy/Downy, Great Spotted/Middle Spotted, Crimson-breasted/Darjeeling? Is this unique for the woodpeckers?
There is at least one theory, which is called "interspecific social dominance mimicry", and was discussed briefly in [this thread].
(Prum R. 2014. Interspecific social dominance mimicry in birds. Zool. J. Lin. Soc. 172: 910-941. [pdf])

(The criticism of the theory in the above thread by Kirk Roth, re. the possibility that the Downy/Hairy similarity may be symplesiomorphic, does not hold is you consider the whole pattern. Eg., the common ancestor of Downy and Hairy is also an ancestor of both Crimson-breasted and Lesser Spotted, but not of Darjeeling or White-backed. If this ancestor was Downy/Hairy-like, it can have been neither Darjeeling-, nor White-backed-like, and a Crimson-breasted/Darjeeling or Lesser Spotted/White-backed similarity can then only be explained by convergence. Quite a few other examples are proposed in the paper.)
 
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There is at least one theory, which is called "interspecific social dominance mimicry", and was discussed briefly in [this thread].
(Prum R. 2014. Interspecific social dominance mimicry in birds. Zool. J. Lin. Soc. 172: 910-941. [pdf])

(The criticism of the theory in the above thread by Kirk Roth, re. the possibility that the Downy/Hairy similarity may be symplesiomorphic, does not hold is you consider the whole pattern. Eg., the common ancestor of Downy and Hairy is also an ancestor of both Crimson-breasted and Lesser Spotted, but not of Darjeeling or White-backed. If this ancestor was Downy/Hairy-like, it can have been neither Darjeeling-, nor White-backed-like, and a Crimson-breasted/Darjeeling or Lesser Spotted/White-backed similarity can then only be explained by convergence. Quite a few other examples are proposed in the paper.)

To use Prum's language from Table 1, I don't see why all four of Downy/Hairy and Lesser Spotted/White-backed aren't "closely similar" (not to mention other species), and thus possibly all sharing a similar-plumaged ancestor. I wrote in the other thread before I had an appreciation for the Crimson-breasted/Darjeeling similarity and admittedly it is uncanny. Perhaps Dryobates is indeed a "mimic genus," but I still contend that without behavioral studies, social mimicry is still just one hypothesis among many to explain plumage convergence in the same habitat/range.
 
Re, Chloropicus:
In another publication from Malherbe in 1845 he lists two species in Picus (Chloropicus) viridis and canus.
https://books.google.com/books?id=V...ce=gbs_ge_summary_r&cad=0#v=onepage&q&f=false . Cahier 3 page 62.

Zoonomen says:
Picus viridis Linnaeus 1758 Syst.Nat.ed.10 p.113
• Type by subsequent designation. Picus viridis Linnaeus (Swainson 1820 Zool.Ill. 1 pl.4text)???

Here is the original description of Picus syriacus:
http://docnum.unistra.fr/cdm/compoundobject/collection/coll13/id/126954/rec/6 .
Signature or Folio r note 5. Page 00043. One author misread the small r as a v. This is the first I have seen Hemprich & Ehrenberg online.
Someone is selling a drawing from this book to profit African nature.
http://www.ebay.com/itm/1829-Ehrenb...560?pt=LH_DefaultDomain_0&hash=item3cbfebc1b0 .
A journal that Bird Forum nomenclature types might like.
http://www.biotaxa.org/sherbornia/issue/view/1220 .
 
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Re, Chloropicus:
In another publication from Malherbe in 1845 he lists two species in Picus (Chloropicus) viridis and canus.
https://books.google.com/books?id=V...ce=gbs_ge_summary_r&cad=0#v=onepage&q&f=false . Cahier 3 page 62.
Interesting--I had missed this one. The "3e cahier" is just dated "1845" [here]. In the work [here], Malherbe refers to "Revue zool. 1845, p. 375.", which is where he described Picus (Leuconotopicus) Numidicus [OD], in the "October 1845" issue of Revue zoologique. The work that is accepted as having introduced Chloropicus was published in the next issue ("November 1845") of the same journal [here].
Should this "3e cahier" happen to have been published between the "October 1845" and the "November 1845" issues of Revue zoologique, the originally included species of Chloropicus would be viridis and canus, and Stejneger's 1886 designation of viridis would be valid...
(It was in any case published before the "February 1846" issue of Revue zoologique, because [there], Malherbe cited this work. Pica Mauritanica Malherbe [OD], which was described in the "3e cahier", is usually cited from 1845.)
Zoonomen says:
Picus viridis Linnaeus 1758 Syst.Nat.ed.10 p.113
• Type by subsequent designation. Picus viridis Linnaeus (Swainson 1820 Zool.Ill. 1 pl.4text)???
This is the type fixation of Picus Linnaeus. Info apparently taken from Peters 1964 [here], and not fully correct: the designation is actually in the text accompanying plate 14, [here]. "Picus rubiginosus" [...] "Generic type Picus viridis." Stresemann 1921 [here] had this right.
Some early authors have considered that the type was Picus major Linnaeus; others that it was Picus martius Linnaeus. In these cases, Gecinus Boie 1830 was most often used for green woodpeckers, as it has precedence over Chloropicus Malherbe 1845.
 
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*sigh* I wish things were more frequently available to those of us who have the interest but not the institutional affiliation. :/
 
*sigh* I wish things were more frequently available to those of us who have the interest but not the institutional affiliation. :/
There's free/open access to all full-text papers posted on ResearchGate – just dismiss the invitation to sign-up if it appears.

PS. There currently seems to be a (temporary?) problem with the direct link to Fuchs & Pons 2015. But the paper is still accessible by clicking on the 'Source' image on Jérôme Fuchs's contributions page.
 
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Perhaps Laurent should have been added to the acknowledgements section...

I guess it would be quicker to route papers via Laurent pre-submission!:t:

On the other hand, we wouldn't have the pleasure in seeing his deconstructing logic at work...:eek!:
MJB
 
Léo Signorini Novaes, 2013. Osteological adaptation for terrestrial habit in woodpeckers (Aves: Picidae). Dissertation, Universidade de São Paulo.

Abstract:
Woodpeckers (family Picidae) are a well-defined group, traditionally characterized by their unique morphological specializations for climbing and excavating in trees, like posterior limbs and the tail adapted for support and a robust cranium and beak for hammering on wood. Among the species in the family, however, there are not only some that stray from this highly specialized behavior and develop ground-based behavior, but also a few that completely abandon trees. The genus with most of these cases, Colaptes Vigors, 1825, contains species that specialize in feeding on ants ant termites and their larvae, building nests in banks, holes in the ground or even termite hills. Besides them, the monotipical African genus Geocolaptes Gmelin, 1788 shows the same feeding and nesting habits but is much more distantly related. This work aims at analyzing relations between the shapes of the species' skeletons and looking for similarities and differences that correlated with their habitats and behavior and whether or not the similar solutions occurred independently throughout the family's evolutionary history. 49 skeleton specimens were utilized, belonging to the two ground dwelling and the Dryocopus and Celeus genera. Principal Component Analysis showed correlation between feeding and nesting habits and several measures of the skull (skull height, beak shape and nostrils' shape and positioning) and postcrania (size and shape of the pygostyle and size of the keel). It is a possibility that these changes in shape derive from two distinct selective pressures: the exploration of a different food source (ants and termites) and the change in the way the organisms move (pygostyle as a support in climbing and the keel relating to the strain of taking off). The results suggest that these adaptations came about independently but generated extremely similar shapes, while also demonstrating the need for further study with the inclusion of more taxa for a more representative comparison
 

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