Also, I wonder why lesser short-toed lark isn't listed in the Birds of Central Asia guidebook.
It may be because the identity, relationships and distributions of populations were less than clear; also some taxa were mis-attributed, if we go by the revisions of Alström
et al 2013 & Stervander
et al 2016,both clarifying many aspects, but after Ayé
et al 2012.
Similar taxa likely present in the Aral region, in post-revision proposed taxonomy, are: Greater Short-toed Lark
Calandrella brachydactyla longipennis, Hume's Short-toed Lark
Calandrella acutirostris acutirostris (east of Aral, mostly), Lesser Short-toed Lark
Alaudala rufescens aharoni (including
niethammeri and Asian Short-toed Lark
Alaudala cheleensis leucophaea (rare resident in and W of Syrdarya).
Because distributions remain poorly-known, other taxa may also be present. Note that previous Russian taxonomy (including unilateral adoption of some English names which in the West have been used for more distant taxa) may yet provide some clarity or even cloud the issue.
MJB
Alström, P, KN Barnes, U Olsson, FK Barker, P Bloomer, AA Khan, MA Quresh, A Guillaumet, P-A Crochet and PG Ryan. 2013. Multilocus phylogeny of the avian family Alaudidae (larks) reveals complex morphological evolution, non-monophyletic genera and hidden species diversity.
Mol. Phyl. & Evol.
69(3): 1043-1056.
Ayé, R, M Schweizer and T Roth. 2012. The Birds of Central Asia. Helm. London. UK.
Stervander, M, P Alström, U Olsson, U Ottosson, B Hansson and S Bensch. 2016. Multiple instances of paraphyletic species and cryptic taxa revealed by mitochondrial and nuclear RAD data for
Calandrella larks (Aves: Alaudidae).
Mol. Phyl. Evol.
102: 233–245