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Kentish Plover (1 Viewer)

Richard Klim

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Küpper et al 2009. Kentish versus Snowy Plover: Phenotypic and Genetic Analyses of Charadrius alexandrinus Reveal Divergence of Eurasian and American Subspecies. The Auk: in press.
http://caliber.ucpress.net/doi/abs/10.1525/auk.2009.08174

Recommends splitting of C nivosus (already recognised by Rasmussen & Anderton 2005, and IOC). Is occidentalis considered? (Also identified as a possible species, 'Peruvian Plover', by Monroe & Sibley 1993.)

Richard
 
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No, they only sampled NA birds of subspecies nivosus. In Eurasia, they only sampled nominate subspecies. They included samples of African White-fronted Plover, and that seems to be closer to alexandrinus than nivosus is. They did not sample far east birds, and that might be a weakness in their study, I am curious to see what SACC and NACC will think about this.

Niels
 
The main author is currently working on the genetics of the White-faced Plover and this should be completed by the end of November.
Is this the putative 'new' plover from south-east Asia.
Yes. Kennerley et al 2008 (Rediscovery of a long-lost Charadrius plover from South-East Asia. Forktail 24) suggests that 'White-faced Plover' is the true Charadrius [alexandrinus] dealbatus, and that the name nihonensis (Deignan, 1941) should apply to the form of Kentish Plover breeding in north-eastern Asia.

Richard
 
BNA Online

A new version (Page et al, 18 Nov 2009) of the BNA Online species account for Charadrius alexandrinus has been posted, which acknowledges recent work:

"The family Charadriidae, the plovers and lapwings, is well supported and is one of the core groups in the shorebird (Charadriiformes) radiation. Within this family, the genus Charadrius is also well defined. A mitochondrial DNA-based phylogeny of thirteen members of the genus (Joseph et al. 1999) found C. alexandrinus to be sister to C. ruficapillus (the Red-capped Plover of Australia), and these two species, along with C. marginatus (the White-fronted Plover of sub-Saharan Africa) and C. javanicus (the Javan Plover of Java), form a superspecies (Wiersma 1996). On the basis of its geographic distribution and morphology, C. peronii (the Malaysian Plover of Indonesia and the Philippines) may also be an allospecies in this superspecies complex.

Species limits within this superspecies are open to debate. For example, Wiersma (1996) said of C. javanicus that it "may well not merit full species status" and noted that it is "often" treated conspecific with C. alexandrinus. At the other extreme, Howard and Moore (1994) treated C. a. occidentalis as a species despite its strong morphological similarity to C. a. nivosus (see also Sibley and Monroe 1990). Species limits of C. a. dealbatus are being explored currently (Kennerley et al. 2008), and it is possible that this taxon will be accord species status once its breeding range and behavior are elucidated fully.

It is also possible that taxa in the Old World and New World will be split from each other, a Kentish Plover vs. Snowy Plover split. Ridgley and Greenfield (2001) remarked that "American birds are perhaps better treated as a distinct species (C. nivosus) separate from those of the Old World; their calls appear to differ," but provided no data in support. Küpper et al. (2009), however, provided data on molecular (mtDNA, a nuclear gene, and 26 microsatellites) divergence, as well as detailing differences in courtship calls and downy plumage of chicks. The genetic analyses demonstrated that C. a. alexandrinus is sister to C. marginatus, not to C. a. nivosus. If true, then the resultant paraphyly likely will lead taxonomists either to lump C. marginatus with the C. alexandrinus complex or split C. alexandrinus into Old World and New World species."​

Richard
 
The genetic analyses demonstrated that C. a. alexandrinus is sister to C. marginatus, not to C. a. nivosus. If true, then the resultant paraphyly likely will lead taxonomists either to lump C. marginatus with the C. alexandrinus complex or split C. alexandrinus into Old World and New World species.

Looking at mtDNA alone (here, DNA barcodes), it doesn't actually seem that lumping marginatus with alexandrinus would suffice to eliminate the paraphyly...
 

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Looking at mtDNA alone (here, DNA barcodes), it doesn't actually seem that lumping marginatus with alexandrinus would suffice to eliminate the paraphyly...
Indeed - pallidus would also need to be lumped. Surely better to split nivosus...

Richard
 
To help play the home version of the game:

…lumped in the 19th Supplement (first Supplement since the Check-List) in 1944 (Auk 61:441-464).
http://elibrary.unm.edu/sora/Auk/v061n03/p0441-p0464a.pdf .

104 Charadrius nivosus nivosus and C. n. tenuirostris become Charadrius alexan-
drinus nivosus and Charadrius alexandrinus tenuirostris, as they are con-
sidered races of the Old World bird. See Peters, Check-list, 2, 1934, 250.

Charadrius nivosus (Cass.).
The bird described by Cassin as Aegialitis nivosa has by most recent authors been considered a distinct species. It is very similar in general appearance to Charadrius alexandrinus alexandrinus (Linnaeus), but differs chiefly in having the lores wholly white (instead of with a black line from the eye to the bill), and the rufous of the pileurn less pronounced in fully adult plumage. Examination of a large series of birds of this species shows, however, that the characters distinguishing these two forms are not constant. That of the rufous of the head is too changeable to be of much value
in the determination of specimens, so that the color of the lores is much the best character. But even this varies to such an extent that it is impossible to find any definite llne of demarcation between the two forms. Specimens of the American Charadrius nivosus occur with partly black lores, and of the Old World Charadrius
alexandrinus with nearly white lores. In view of these facts, it is of course, inadvisable to maintain the specific distinction of Charadrius nivosus. It, therefore, should be regarded as a subspecies of Charadrius alexandrinus. It is interesting to note that this is the view taken many years ago by Mr. Henry Seebohm, as well as by some recent authors. Mr. Ridgway has recently separated the American birds into two races; so that our Snowy Plovers should now stand as Charadrius alexandrinus nivosus
(Cassin), and Charadrius alexandrinus tenuirostris (Lawrence).
1 Rep. Explor. and Surv. R. R. Pac., IX, 1858, p. 696 (Presidio, near San
Francisco, California).
2 Geographical Distribution of the Charadriidae, 1887, p. 171.

3 Bull. U.S. Nat. Mus., No. 50, part VIII, 1919, pp. 136-140

Leucopolius alexandrinus nivosus (Cassin) becomes Charadrius
nivosus nivosus (Cassin), because specifically different from Cha-
radrius nivosus, and not generically distinct from Charadrius. (Cf.
Ridgway, Bull. U.S. Nat. Mus., No. 50, VIII, June 26, 1919, pp.
136-137.)
Charadrius nivosus tenuirostris (Lawrence). Aegialitis tenuirostris
Lawrence, Ann. Lyc. Nat. Hist. N.Y., VII, 1862, p. 455 (near Guan-
tanamo, Cuba). Revived as a subspecies. (Cf. Ridgway, Bull.
U.S. Nat. Mus., No. 50, VIII, June 26, 1919, p. 139.) Range:
southeastern United States, south in winter to Paraguay.

FIFTH ANNUAL LIST OF PROPOSED CHANGES IN THE
A. O. U. CHECK-LIST OF NORTH AMERICAN BIRDS.
BY HARRY C. OBERHOLSER. Auk April 1920.

Peters Birds of the World v. 2
http://www.biodiversitylibrary.org/item/50592#page/272/mode/1up .




Linnaeus, C. 1758. Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 10th edition.


http://www.biodiversitylibrary.org/item/10277#page/169/mode/1up .
Charadrius alexandrinus habitat ad Aegypteus ex Nilo canalem.
He takes the name from Hasselquist. The Kent Plover name comes from a Latham name from 1790 (Charadrius cantianus), so the vernacular name should be either Alexandrian Plover, Nile Channel Plover, or Hasselquist’s Plover. Charadrius alexandrinus m. The Alexandrian Dotterell. IT is of the size of a Lark: I found it the 24th of May, 1750, in the channel, which leads the water of the Nile to Alexandria. It feeds on insects.
http://books.google.com/books?pg=PA...Zwd2_Cw&ct=result&id=8lFCAAAAcAAJ&output=text .



…tenuirostris
http://books.google.com/books?id=Vg...&resnum=2&ved=0CB4Q6AEwAQ#v=onepage&q&f=false .
p. 455.
His first visit to the rich collecting grounds about Guantanamo was in June, 1858, when he lived six weeks at the railway station at Caimanera on Guantanamo Bay, where he collected along its shores. Having found in July, on one of the immense sand flats of that region, a nesting Snowy Plover, Aegialitis nivosa (Cassin) with a set of three eggs, he took the mother alive in a bird-net, sending her to Mr. Lawrence, who described it as Aegialitis tenuirostris (Law.), but Dr. Ridgway later determined the species to be A. nivosa (Cass.).

Occidentalis
Charadrius nivosus occidentalis (Cabanis) 1872 J.Orn. 20 no.116 p.158 .
I think occidentalis came from Peru? Chile? Not a “North American subspecies”.

Charadrius marginatus Vieillot 1818 Nouv.Dict.Hist.Nat. 27 p.138 .
In this Vieillot says Charadrius marginatus is from Geoffrey de Saint-Hilaire. Zoonomen has this from the Dictionary not Ornithologie
http://books.google.com/books?q=editions:LCCN06002713&lr=&id=_1JhAAAAIAAJ&sa=N&start=40 . Volume 27 p. 138.

Measurements of C. alexandrinus (Charadrius alexandrinus Linnaeus; C. cantianut Latham) are given for comparison, this being the Old World representative of C. nivosus. Besides having longer wings, relatively shorter tail, and decidedly longer tarsi and toes than C. nivosus, C. alexandrinus has a black loral stripe and other differences of coloration.
Ridgway 1919
The birds of North and middle America
http://books.google.com/books?print...&pg=PA137&id=5ooaAAAAYAAJ#v=onepage&q&f=false .

“With a decent sample of published sonograms, I would instantly change to a YES vote,” Someone needs to get to work.
 
Can anyone point me in the direction of the old world subspecies of Kentish plover. (i.e. not Snowy plover sspp)

Trying to find scientific names, locations, differences

Thanks
 
Subspecies

eg, Wiersma 1996 (HBW 3).

BWP:
Nominate alexandrinus Linnaeus, 1758, Eurasia and northern Africa, south to Mauritania, Gulf of Aden, Socotra, and western India, east to south-east Transbaykalia (USSR) and Inner Mongolia. Extralimital: dealbatus (Swinhoe, 1870), Japan, eastern China, and Ryukyu Islands, intergrading with nominate alexandrinus in Manchuria, Korea, and southern Ussuriland; nivosus (Cassin, 1858), western and central USA; tenuirostris (Lawrence, 1862), south-east USA and Caribbean; occidentalis (Cabanis, 1872), coastal western South America; seebohmi Hartert and Jackson, 1915, Ceylon.

Geographical variation
Mainly clinal in nominate alexandrinus; too much overlap in characters to split this race further, as sometimes advocated. Length of wing, tail, and toe gradually decreases from western and southern Europe through North Africa towards southern Red Sea and Gulf of Aden; variation in bill and tarsus slight, not gradually clinal. Birds of Persian Gulf and north-east Arabia also small; those breeding inland Iraq and Iran of same size as North African birds. Slight clinal increase in wing from southern Europe eastward to central Asia; average wing of adult ♂♂ from Sinkiang, Tibet, and Mongolia 113.4–114 (Piechocki 1968; Vaurie 1972; ZMA). Variation in colour slight; adult ♂♂ of southern and eastern populations tend to have more often bright cinnamon crown than those of western Europe, and adult ♀♀ more often tinged cinnamon on upperparts and sides of chest (occasionally central chest also). Upperparts rather similar throughout, but southern populations more strongly subject to bleaching, especially those of Mauritania, Libya, Red Sea, and Persian Gulf, appearing nearly white when worn. East Palearctic dealbatus similar in size and colour to central Asiatic populations of nominate alexandrinus, but bill longer and heavier – length 17.4 (10) 16.8–18.4 and depth at gonys 3.95 (10) 3.6–4.3, against 15.3(110)13.7–17.1 and 3.44 (55) 3.2–3.8 in nominate alexandrinus from Europe and central Asia; dealbatus intergrades with nominate alexandrinus in Manchuria, Korea, and southern Ussuriland (USSR). North American race nivosus with its close Caribbean and South American allies tenuirostris and occidentalis differs from nominate alexandrinus by absence of black line on lores, paler colour of upperparts, and small size; seebohmi, of Ceylon, small and indistinctly patterned.
But, if the recently rediscovered 'White-faced Plover' assumes the name dealbatus, then nihonensis Blakiston 1876 is available for the NE Asian form:
  • Kennerley, Bakewell & Round 2008. Rediscovery of a long-lost Charadrius plover from South-East Asia. Forktail 24: 63-79. [pdf]
 
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I formed an opinion at the time that it was a US bird, based on similarity the Cheshire bird but I only saw it a couple of times, instead of most days for a 7 winters, including in full summer plumage

Ihave early and primitive notes suggesting it was a Snowy.
 

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