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Difference between revisions of "Audubon's Shearwater" - BirdForum Opus

(Corrected small errors and typo in Habitat and Breeding)
(taxonomy (I hope it's clearer now...... ;-) ))
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[[Great Shearwater]] is much larger with lighter body, smudgy brown belly, neat cap, narrow pointed wings. and white tail band.
 
[[Great Shearwater]] is much larger with lighter body, smudgy brown belly, neat cap, narrow pointed wings. and white tail band.
 
==Distribution==
 
==Distribution==
Breeds in the Caribbean. All breeding grounds are islands.
+
Breeds in the Caribbean. All breeding grounds are islands.<br />
 +
Recently discovered off [[Brazil]] on Fernando do Noronha, Itataia Islands.<br />
 +
Winters in the Carribbean and up the east coast of the [[United States]].
 
==Taxonomy==
 
==Taxonomy==
[[Heinroth's Shearwater]], [[Persian Shearwater]], [[Tropical Shearwater]] and [[Galapagos Shearwater]] have been split from Audubon's Shearwater.
+
Clements<sup>[[#References|[1]]]</sup> and Gill and Donsker<sup>[[#References|[3]]]</sup> agree on the taxonomy of this species. Dickinson<sup>[[#References|[4]]]</sup> however treats the shearwaters differently.<br />
 +
[[Tropical Shearwater]] and [[Persian Shearwater]] have in the past been included in this species. Both are now split (however [[Persian Shearwater]] is included in Tropical by Dickinson). The split of [[Galapagos Shearwater]], which is known to differ strikingly both in DNA (with high support), physiology and voice is accepted by all three authorities, the same goes for [[Heinroth's Shearwater]].<br />
 +
Dickinson follows the Austin et al. 2004 revisions of the Audubon's/Little Shearwater complex and has transferred the two subspecies ''baroli'' and ''boydi'' of [[Little Shearwater]] to this species. Clements and Gill and Donsker follow Sangster et al 2005 and Onley & Scofield 2007 in this case and accept both former subspecies as full species ([[Barolo Shearwater]] and [[Boyd's Shearwater]]).
 
====Subspecies====
 
====Subspecies====
Audubon's Shearwater as treated here has only 2 subspecies<sup>[[#References|[1]]]</sup>:
+
Audubon's Shearwater as treated here has 2 subspecies<sup>[[#References|[1]]]</sup>:
 
 
 
*''P. l. lherminieri'':
 
*''P. l. lherminieri'':
 
:*Breeds [[Bahamas]] and [[West Indies]]; formerly [[Bermuda]]
 
:*Breeds [[Bahamas]] and [[West Indies]]; formerly [[Bermuda]]
Line 38: Line 41:
 
:*Islets in south-western [[Caribbean]]
 
:*Islets in south-western [[Caribbean]]
  
Up to 8 subspecies are recognised by various authorities and some of these are treated in detail below:
+
The described subspecies ''atrodorsalis'' (Mascarene Shearwater) is now believed to have been a juvenile of the nominate form of [[Tropical Shearwater]].<br />
====Status of [[Galapagos Shearwater]]====
+
Carboneras et al<sup>[[#References|[2]]]</sup> don't accept ''loyemilleri'' and treat it as a synonym of ''lherminieri''.
According to Boegh (2007), the [[Galapagos Shearwater]] is "now known to differ strikingly both in DNA (with high support), physiology and voice from the members of the Audubon's/Little complex.  Indeed, they're not even the closest relatives of the Galapagos Shearwater, which actually is closer to the [[Christmas Shearwater]]." SACC (2008) treats ''subalaris'' as a full species, citing the genetic data of Austin et al (2004) and long-ago noted morphological distinctiveness by Murphy (1927). However, Sibley & Monroe and Howard & Moore (2003) retain it as a subspecies of ''lherminieri''.
 
====Status of Bannerman's Shearwater====
 
Austin et al (2004) treat ''bannermani'' as a full species based on molecular phylogeny, and are followed by Onley & Scofield (2007). Sibley & Monroe (1996) also grant it species status. However, Howard & Moore (2003) retain it as a subspecies of ''lherminieri'' while the 2009 update to Clements moved the subspecies ''bannermani'' into what they designate to be a newly elevated taxon, ''P. bailloni'', [[Tropical Shearwater]] (see below). The Opus places it into ''P. bailloni''.
 
====Status of [[Tropical Shearwater]]====
 
Austin and others in 2004 in a paper proposing a major rearrangement of taxonomy of smaller shearwaters, proposed that the taxons from tropical Indian and Pacific Oceans should be separated out as [[Tropical Shearwater]], ''Puffinus bailloni''. This taxon includes subspecies ''bailloni'', ''dichrous'' (current subspecies ''nicolae'', ''colstoni'', and ''polynesia'' would disappear into this one), ''temptator'' and ''persicus'' (this latter form is currently considered a full species, [[Persian Shearwater]] by Opus based on a majority of the major checklists). The 2009 update to Clements did in fact elevate ''bailloni'' to species status, including subspecies ''bailloni'', ''dichrous'', ''nicolae'', ''gunax'', and ''bannermani''.
 
====Mascarene Shearwater, ''Puffinus atrodorsalis''====
 
Invalid taxon. Believed to have been a juvenile ''P. b. bailloni'' (Austin et al., 2004).
 
====Other notes====
 
Subspecies ''temptator'' is now believed to belong to [[Persian Shearwater]] but has in the past been included here.
 
 
 
If the Austin et al. 2004 revisions are followed, there would be a transfer of two subspecies from [[Little Shearwater]] to here: subspecies ''baroli'' and ''boydi''. Some suthorities have expected that ''baroli'' and ''boydi'' would become one full species (Macaronesian Shearwater) or two species, a solution followed by Clements 2009.
 
 
==Habitat==
 
==Habitat==
 
Mainly occurs in offshore waters. Near land next to colonies during the breeding season where it breeds on islands, offshore islets and coral atolls, utilising cliffs and slopes with herbaceous vegetation and rocky areas.
 
Mainly occurs in offshore waters. Near land next to colonies during the breeding season where it breeds on islands, offshore islets and coral atolls, utilising cliffs and slopes with herbaceous vegetation and rocky areas.
Line 65: Line 57:
 
Similar to [[Manx Shearwater]] but more strangled. Female calls are lower and harsher than males. Only calls at night. Mainly silent at sea.
 
Similar to [[Manx Shearwater]] but more strangled. Female calls are lower and harsher than males. Only calls at night. Mainly silent at sea.
 
==References==
 
==References==
#{{Ref-Clements6thAug13}}#Handbook of the Birds of the World Alive (retrieved September 2015)
+
#{{Ref-Clements6thAug15}}#Handbook of the Birds of the World Alive (retrieved September 2015)#{{Ref-GillDonsker15V5.3}}#{{Ref-HM04}}
 
#Wikipedia
 
#Wikipedia
 
# [http://www.bioone.org/doi/abs/10.1642/0004-8038%282004%29121%5B0847%3AAGMPOT%5D2.0.CO%3B2 Paper ] by Austin et al. (2004) describing phylogeny of small shearwaters
 
# [http://www.bioone.org/doi/abs/10.1642/0004-8038%282004%29121%5B0847%3AAGMPOT%5D2.0.CO%3B2 Paper ] by Austin et al. (2004) describing phylogeny of small shearwaters

Revision as of 15:36, 1 October 2015

Photo by Pauhana
gulf stream off Florida, September 2014
Puffinus lherminieri

Identification

27–33 cm, 10.6-13 inches. Wingspan 64–74 cm, 25-29 inches. 138–290 g

  • Blackish-brown upperparts.
  • Underparts, cheeks and throat are white with conspicuos spot at sides of rump.
  • Wing are broad and short.
  • Black tail is long and rounded.
  • Black flight feathers.
  • White around eye in most birds.
  • Partial eye ring
  • Occasionally short supercilium in front of eye.
  • Occasionally short and narrow eye stripe behind eye.
  • Dark flanks merge into white breast and occasionally with partial breast band.
  • Thighs and undertail coverts dark brown.
  • Dark grey trailing edge to underwing.
  • Base of primaries normally have small quantity of silver or white
  • Lesser coverts darkly spotted or streaked.
  • Iris dark brown.
  • Beak grey to blue grey often darker on culmen and black tip.
  • Legs and feet pale pink with outer tarsi and toes blackish.
  • Sexes similar, female slightly smaller.

Similar Species

Manx Shearwater is slightly larger with longer more-pointed wings, and white undertail coverts.
Cory's Shearwater is much larger with much lighter brown body and wings, heavy yellow bill and white tail band.
Great Shearwater is much larger with lighter body, smudgy brown belly, neat cap, narrow pointed wings. and white tail band.

Distribution

Breeds in the Caribbean. All breeding grounds are islands.
Recently discovered off Brazil on Fernando do Noronha, Itataia Islands.
Winters in the Carribbean and up the east coast of the United States.

Taxonomy

Clements[1] and Gill and Donsker[3] agree on the taxonomy of this species. Dickinson[4] however treats the shearwaters differently.
Tropical Shearwater and Persian Shearwater have in the past been included in this species. Both are now split (however Persian Shearwater is included in Tropical by Dickinson). The split of Galapagos Shearwater, which is known to differ strikingly both in DNA (with high support), physiology and voice is accepted by all three authorities, the same goes for Heinroth's Shearwater.
Dickinson follows the Austin et al. 2004 revisions of the Audubon's/Little Shearwater complex and has transferred the two subspecies baroli and boydi of Little Shearwater to this species. Clements and Gill and Donsker follow Sangster et al 2005 and Onley & Scofield 2007 in this case and accept both former subspecies as full species (Barolo Shearwater and Boyd's Shearwater).

Subspecies

Audubon's Shearwater as treated here has 2 subspecies[1]:

  • P. l. lherminieri:
  • P. l. loyemilleri:

The described subspecies atrodorsalis (Mascarene Shearwater) is now believed to have been a juvenile of the nominate form of Tropical Shearwater.
Carboneras et al[2] don't accept loyemilleri and treat it as a synonym of lherminieri.

Habitat

Mainly occurs in offshore waters. Near land next to colonies during the breeding season where it breeds on islands, offshore islets and coral atolls, utilising cliffs and slopes with herbaceous vegetation and rocky areas.

Behaviour

Flight

In calmer winds this is a species that tends to fly low. However, if going across strong winds, a high arching flight may be seen that reminds of Petrels.

Breeding

It is a colonial breeder, breeding frequently in low numbers occasionally in mixed colonies, nesting in small burrows and crevices in rocks. Colonies are visited on up to 95% of nights after brooding for 3-7 days, but may be seen from shore during day. The single white egg is incubated by both parents for 44-60 days in stints of between 2-10 days; the young fledge between 62 and 100 days later. Breeding season length and time is dependent on location. Mainly in January to March. In higher latitudes from May to July. Brazilian populations lay eggs between August and September, fledging in October. Sexual maturity is reached at about 8 years. One ringed bird is known to have reached 19 years.

Diet

The diet consists of small fish, squid and crustaceans. They feed by pursuit diving and plunging, and 'pattering' on the water surface and surface grabbing. Dives last up to several seconds. Dives up to 15 m. May join other feeding seabirds and small fishing vessels.

Vocalisation

Similar to Manx Shearwater but more strangled. Female calls are lower and harsher than males. Only calls at night. Mainly silent at sea.

References

  1. Clements, J. F., T. S. Schulenberg, M. J. Iliff, D. Roberson, T. A. Fredericks, B. L. Sullivan, and C. L. Wood. 2015. The eBird/Clements checklist of birds of the world: v2015, with updates to August 2015. Downloaded from http://www.birds.cornell.edu/clementschecklist/download/
  2. Handbook of the Birds of the World Alive (retrieved September 2015)#Gill, F and D Donsker (Eds). 2015. IOC World Bird Names (version 5.3). Available at http://www.worldbirdnames.org/.
  3. Dickinson, EC, ed. 2014. The Howard and Moore Complete Checklist of the Birds of the World. 4th ed. Princeton: Princeton Univ. Press. ISBN 978-0956861122
  1. Wikipedia
  2. Paper by Austin et al. (2004) describing phylogeny of small shearwaters

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