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Cyclopsitta Reichenbach, 1850, versus Opopsitta P. Sclater, 1860. (1 Viewer)

James Jobling

Well-known member
England
I am revisiting these names. Both Howard & Moore, ed. 4, 2013, Vol. 1, p. 383, and HBW Illustrated Checklist, 2014, Vol. 1, p. 748, recognise Cyclopsitta Reichenbach, 1850, for the small Australasian fig parrots. Both works include Opopsitta P. Sclater, 1860, as a synonym of Cyclopsitta, and refer to R. Schodde, 1978, 'The identity of five type-specimens of New Guinean birds', The Emu, 78 (1), 1-6. I agree with Mathews 1912 who rejects Cyclopsitta because the figures shown on Reichenbach's plate LXXXII show composite details; he is supported by Peters' Check-list, 1937, Vol. III, p. 164, where Opopsitta Sclater is used, with a note,"Replaces Cyclopsittacus "Reichenbach," i.e. Sundevall, of Sharpe's Hand-list; name rejected by Mathews as indeterminable (Nov. Zool., 18, 1912, p. 261)" (see Cyclopsitta entry in HBWAlive Key). However, I have yet to see Schodde's 1978 paper, but would like to read it before proceeding further. BHL do not show The Emu for that year, so can any reader or enthusiast oblige?
 
James,

I know the argument advanced by Mathews is of a kind that has been used a lot in ornithology, but the present Code does not allow to reject an available genus-group name based for this type of reason.

What determines the application of a genus-group name is its type species. This type species must be one of the 'originally included nominal species', which are the nominal species included in the genus and cited by an available species-group name in the original description (see [Art. 67.2 of the ICZN]). If no included species is cited by an available name in the OD (as is the case here), you are not supposed to try to ID what appears on the plate. You are supposed to look for the first subsequent work where one or more species were expressly included in the genus and cited by an available name ([67.2.2]): these will act as the originally-included nominal species, whatever the actual ID of the illustrated bird. If there is only one, it is the type species by subsequent monotypy ([69.3]); if there is more, the type species needs to be fixed by a subsequent designation of one of the first subsequently included nominal species ([69.1]). What determines the identity of this type species is the type material associated to the available name by which it is cited; not what appeared on the illustration that made the generic name available.

Cyclopsitta Reichenbach 1850 [OD] was made available by an accompanying illustration, without any included nominal species. Three year later, Jacquinot & Pucheran 1853 [here] then combined Cyclopsitta with the available species-group name diophthalma Hombron & Jacquinot 1841 [OD]: Psittacula diophthalma Hombron & Jacquinot 1841 acts as the only originally included nominal species, and is the type species by subsequent monotypy.

A type species fixed by this method is validly fixed even if it denotes something else than what appeared on the plate. Should the plate demonstrably show another species, your only liberty would be to argue that the type species was misidentified, and to publish an act under [Art. 70.3], which would allow you to either confirm that the fixed nominal species is the indeed type, even though it is not what appeared on the plate, or to 'correct' the identification by designating 'the taxonomic species actually involved in the misidentification' ([70.3.2]). Rejecting the type fixation is not an option; neither is rejecting the name as a whole.

Under the present rules, the only way that a genus-group name could ever really be unidentifiable is if a nomen dubium becomes validly fixed as its type species.
 
Many thanks, Mark, for the link to Schodde's article in NLA News, 2000, X (5), including a précis of Schodde 1978, which was especially enlightening. Reichenbach 1850 obviously based his figures on Plate 25Bis of the Zoology plates in Voyage Pole Sud (Jacquinot & Pucheran 1853). The head is not very good, but figure B, showing the ridged lower mandible, is the clincher which Mathews 1912 overlooked. I shall retain the HBWAlive Key entries suitably modified.
Thanks also, Laurent, for calmly explaining the bureaucratic nightmare that is the current ICZN .
 
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