Thanks for those links - such beautiful creatures - but numbers are not increasing and seem under massive pressure ..... I can't believe they are still hunted. People are so stupid.
I seem to recall that the doco I watched had cougars with radio collars on - so perhaps that earlier study - they were in a snow covered environment. They had designations such as F3, F4 etc. Their survival rates then were below replacement rates. I seem to recall that the successful breeding female was killed (by a male cougar? or wolves? - I can't remember). They were constantly driven off their kills, and I think just about all the cubs died within the first year.