Estrildidae (1 Viewer)

Microscelis amaurotis matchie Momiyama.
In 2002 Dickinson and Morioka et al said about the name:
M.D. Bruce (in litt.) kindly advised us that this emendation was authored by Momiyama (1928). HM reports that Momiyama reverted to matchie in later work and the suggested emendation although technically an accurate rendering may have simply been a lapsus. Momiyami 1928 is Momiyama, T.T., 1928. Twelve forms of Japanese birds. Annot. Orn. Orient. 1: 419-456..See here on Google Books:
https://books.google.com/books?id=Bc1FAAAAYAAJ&focus=searchwithinvolume&q=matchie .
Momiyama uses both spellings on page 430.
Dickinson & Morioka et al. in 2005 said something different.
This name was emended to matchiae by Momiyama
(1940:44).This emendation is too late to be accepted and does not meet the
requirements of Art, 32.5.1,1,(ICZN,l999).
Momiyama 1940 is Avifauna hatiziosimensis. On the resident birds of Hatizyo-zima, southern Seven Islands. Kagaku-no-Nogyo, 20(3/4): 31-44.
The bird was named for Momiyama's Mother and he forgot an 'a'. Most Mothers are female and any bird named for them should end in ae. Bjorn has shown that many Doubutsugaku zasshi are online.
http://ci.nii.ac.jp/vol_issue/nels/AN00166645_en.html .
But not this one.

Urban Olsson, Per Alström. A comprehensive phylogeny and taxonomic evaluation of the waxbills (Aves: Estrildidae). Molecular Phylogenetics and Evolution. In Press, Journal Pre-proof, Available online 3 February 2020. https://doi.org/10.1016/j.ympev.2020.106757

Abstract:

We present a revised taxonomy of Estrildidae based on the first time-calibrated phylogeny of the family Estrildidae estimated from a data set including the majority of the species, and all genera except the monospecific Paludipasser, using two mitochondrial and five nuclear markers. We find that most differences in current taxonomy reflect alternative opinions among authors regarding inclusiveness of genera, which are usually not in conflict with the phylogeny. The most notable exception is the current circumscriptions of the genera Neochmia, Nesocharis and Taeniopygia, which are incompatible with the phylogeny.

Estrildidae is subdivided into six well supported subclades, which we propose be recognized as the subfamilies Amandavinae, Erythrurinae, Estrildinae, Lagonostictinae, Lonchurinae and Poephilinae.

My new classification of Estrildidae based on their result:

Family Estrildidae Bonaparte, 1850
Subfamily Estrildinae Bonaparte, 1850

Genus Brunhilda Reichenbach, 1862
Brunhilda charmosyna (Reichenow, 1881)
Brunhilda erythronotos (Vieillot, 1817)

Genus Coccopygia Reichenbach, 1863
Coccopygia bocagei Shelley, 1903
Coccopygia melanotis (Temminck, 1823)
Coccopygia quartinia (Bonaparte, 1850)

Genus Cryptospiza Salvadori, 1884
Cryptospiza jacksoni Sharpe, 1902
Cryptospiza reichenowii (Hartlaub, 1874)
Cryptospiza salvadorii Reichenow, 1892
Cryptospiza shelleyi Sharpe, 1902

Genus Delacourella Wolters, 1949
Delacourella capistrata (Hartlaub, 1861)

Genus Estrilda Swainson, 1827
Estrilda astrild (Linnaeus, 1758)
Estrilda atricapilla Verreaux & Verreaux, 1851
Estrilda kandti Reichenow, 1902
Estrilda melpoda (Vieillot, 1817)
Estrilda nigriloris Chapin, 1928
Estrilda nonnula Hartlaub, 1883
Estrilda ochrogaster Salvadori, 1897
Estrilda paludicola von Heuglin, 1863
Estrilda poliopareia Reichenow, 1902
Estrilda rhodopyga Sundevall, 1850
Estrilda rufibarba (Cabanis, 1851)
Estrilda troglodytes (Lichtenstein, 1823)

Genus Glaucestrilda Roberts, 1922
Glaucestrilda caerulescens (Vieillot, 1871)
Glaucestrilda perreini (Vieillot, 1817)
Glaucestrilda thomensis (de Sousa, 1888)

Genus Mandingoa Hartert, 1919
Mandingoa nitidula (Hartlaub, 1865)

Genus Nesocharis Alexander, 1903
Nesocharis ansorgei (Hartert, 1899)
Nesocharis shelleyi Alexander, 1903

Genus Nigrita Strickland, 1843
Nigrita bicolor (Hartlaub, 1844)
Nigrita canicapillus (Strickland, 1841)
Nigrita fusconotus Fraser, 1843
Nigrita luteifrons Verreaux & Verreaux, 1851

Genus Parmoptila Cassin, 1859
Parmoptila jamesoni (Shelley, 1890)
Parmoptila rubrifrons (Sharpe & Ussher, 1872)
Parmoptila woodhousei Cassin, 1859

Subfamily Lagonostictinae Steiner, 1960

Genus Clytospiza Shelley, 1896
Clytospiza monteiri (Hartlaub, 1860)

Genus Euschistospiza Wolters, 1943
Euschistospiza cinereovinacea (de Sousa, 1889)
Euschistospiza dybowskii (Oustalet, 1892)

Genus Granatina Sharpe, 1890
Granatina granatina (Linnaeus, 1766)
Granatina ianthinogaster (Reichenow, 1879)

Genus Hypargos Reichenbach, 1862
Hypargos margaritatus (Strickland, 1844)
Hypargos niveoguttatus (Peters, 1868)

Genus Lagonosticta Cabanis, 1851
– Subgenus Hypargoides Wolters, 1943
Lagonosticta (Hypargoides) nitidula Hartlaub, 1886
Lagonosticta (Hypargoides) rufopicta (Fraser, 1843)
Lagonosticta (Hypargoides) senegala (Linnaeus, 1766)
– Subgenus Lagonosticta Cabanis, 1851
Lagonosticta (Lagonosticta) landanae Sharpe, 1890
Lagonosticta (Lagonosticta) rhodopareia (von Heuglin, 1868)
Lagonosticta (Lagonosticta) rubricata (Lichtenstein, 1823)
Lagonosticta (Lagonosticta) sanguinodorsalis Payne, 1998
Lagonosticta (Lagonosticta) umbrinodorsalis Reichenow, 1910
Lagonosticta (Lagonosticta) virata Bates, 1932
– Subgenus Lychnidopsiza Heuglin, 1871
Lagonosticta (Lychnidospiza) rara (Antinori, 1864)
– Subgenus Mormolycia Reichenbach, 1862
Lagonosticta (Mormolycia) larvata (Rüppell, 1840)
Lagonosticta (Mormolycia) nigricollis Heuglin, 1863
Lagonosticta (Mormolycia) vinacea (Hartlaub, 1857)

Genus Pyrenestes Swainson, 1837
Pyrenestes minor Shelley, 1894
Pyrenestes ostrinus (Vieillot, 1805)
Pyrenestes sanguineus Swainson, 1837

Genus Pytilia Swainson, 1837
Pytilia afra (Gmelin, 1789)
Pytilia hypogrammica Sharpe, 1870
Pytilia lineata von Heuglin, 1863
Pytilia phoenicoptera Swainson, 1837

Genus Spermophaga Swainson, 1837
Spermophaga haematina (Vieillot, 1808)
Spermophaga poliogenys (Ogilvie-Grant, 1906)
Spermophaga ruficapilla (Shelley, 1888)

Genus Uraeginthus Cabanis, 1851
Uraeginthus angolensis (Linnaeus, 1758)
Uraeginthus bengalus (Linnaeus, 1766)
Uraeginthus cyanocephalus (Richmond, 1897)

Genus Zonogastris Cabanis, 1851
Zonogastris melba (Linnaeus, 1758)

Sous-famille Amandavinae Steiner, 1960

Genus Amadina Swainson, 1827
Amadina erythrocephala (Linnaeus, 1758)
Amadina fasciata (Gmelin, 1789)

Genus Amandava Blyth, 1836
Amandava amandava (Linnaeus, 1758)

Genus Ortygospiza Sundevall, 1850
Ortygospiza atricollis (Vieillot, 1817)
Ortygospiza fuscocrissa von Heuglin, 1863
Ortygospiza gabonensis Lynes, 1914

Genus Paludipasser Neave, 1909
Paludipasser locustella Neave, 1909

Genus Sporaeginthus Cabanis, 1851
Sporaeginthus subflavus (Vieillot, 1819)

Genus Stictospiza Sharpe, 1890
Stictospiza formosa (Lathamn, 1790)

Subfamily Erythrurinae Delacour, 1943

Genus Chloebia Reichenbach, 1862
Chloebia gouldiae (Gould, 1844)

Genus Erythrura Swainson, 1837
– Subgenus Amblynura Reichenbach, 1862
Erythrura (Amblynura) coloria Ripley & Rabor, 1961
Erythrura (Amblynura) cyanovirens (Peale, 1848)
Erythrura (Amblynura) papuana Rothschild & Hartert, 1900
Erythrura (Amblynura) pealii Hartlaub, 1852
Erythrura (Amblynura) psittacea (Gmelin, 1789)
Erythrura (Amblynura) regia (Sclater, 1881)
Erythrura (Amblynura) tricolor (Vieillot, 1817)
Erythrura (Amblynura) trichroa (von Kittlitz, 1833)
– Subgenus Erythrura Swainson, 1837
Erythrura (Erythrura) prasina (Sparrman, 1788)
Erythrura (Erythrura) viridifacies Hachisuka & Delacour, 1937
– Subgenus Reichenowia Poche, 1904
Erythrura (Reichenowia) hyperythra (Reichenbach, 1862)
– Subgenus Rhamphostruthus Mayr, 1931
Erythrura (Rhamphostruthus) kleinschmidti (Finsch, 1878)

Subfamily Lonchurinae Steiner, 1960

Genus Euodice Reichenbach, 1862
Euodice cantans (Gmelin, 1789)
Euodice malabarica (Linnaeus, 1758)

Genus Lepidopygia Reichenbach, 1862
Lepidopygia nana (Pucheran, 1845)

Genus Lonchura Sykes, 1832
– Subgenus Lonchura Sykes, 1832
Lonchura (Lonchura) punctulata (Linnaeus, 1758)
– Subgenus Munia Hodgson, 1836
Lonchura (Munia) atricapilla (Vieillot, 1807)
Lonchura (Munia) caniceps (Salvadori, 1876)
Lonchura (Munia) castaneothorax (Gould, 1837)
Lonchura (Munia) ferruginosa (Sparrman, 1789)
Lonchura (Munia) flaviprymna (Gould, 1845)
Lonchura (Munia) forbesi (Sclater, 1879)
Lonchura (Munia) grandis (Sharpe, 1882)
Lonchura (Munia) hunsteini (Finsch, 1886)
Lonchura (Munia) maja (Linnaeus, 1766)
Lonchura (Munia) malacca (Linnaeus, 1766)
Lonchura (Munia) melaena (Sclater, 1880)
Lonchura (Munia) montana Junge, 1939
Lonchura (Munia) monticola (De Vis, 1897)
Lonchura (Munia) nevermanni Stresemann, 1934
Lonchura (Munia) pallida (Wallace, 1864)
Lonchura (Munia) quinticolor (Vieillot, 1807)
Lonchura (Munia) spectabilis (Sclater, 1878)
Lonchura (Munia) stygia Stresemann, 1934
Lonchura (Munia) teerinki Rand, 1940
Lonchura (Munia) vana (Hartert, 1930)
– Subgenus Trichogrammoptila Reichenbach, 1862
Lonchura (Trichogrammoptila) fuscans (Cassin, 1852)
Lonchura (Trichogrammoptila) kelaarti (Jerdon, 1863)
Lonchura (Trichogrammoptila) leucogastra (Blyth, 1846)
Lonchura (Trichogrammoptila) leucogastroides (Moore, 1858)
Lonchura (Trichogrammoptila) molucca (Linnaeus, 1758)
Lonchura (Trichogrammoptila) striata (Linnaeus, 1766)

Genus Mayrimunia Wolters, 1949
Mayrimunia leucostista (d'Albertis & Salvadori, 1879)
Mayrimunia tristissima (Wallace, 1865)

Genus Odontospiza Oberholser, 1905
Odontospiza griseicapilla (Delacour, 1943)

Genus Padda Bonaparte, 1850
Padda fuscata (Vieillot, 1807)
Padda oryzivora (Linnaeus, 1758)

Genus Spermestes Swainson, 1837
Spermestes bicolor (Fraser, 1843)
Spermestes cucullata Swainson, 1837
Spermestes fringilloides (Lafresnaye, 1835)

Subfamily Poephilinae Mayr, Paynter & Taylor, 1968

Genus Aegintha Cabanis, 1851
Aegintha temporalis (Latham, 1801)

Genus Aidemosyne Reichenbach, 1862
Aidemosyne modesta (Gould, 1837)

Genus Bathilda Reichenbach, 1862
Bathilda ruficauda (Gould, 1837)

Genus Emblema Gould, 1842
Emblema pictum Gould, 1842

Genus Heteromunia Mathews, 1913
Heteromunia pectoralis (Gould, 1841)

Genus Neochmia Gray, 1849
Neochmia evangelinae d'Albertis & Salvadori, 1879
Neochmia phaeton (Hombron & Jacquinot, 1841)

Genus Oreostruthus De Vis, 1898
Oreostruthus fuliginosus (De Vis, 1897)

Genus Poephila Gould, 1842
Poephila acuticauda (Gould, 1840)
Poephila cincta (Gould, 1837)
Poephila personata Gould, 1842

Genus Stagonopleura Reichenbach, 1850
Stagonopleura guttata (Shaw, 1796)

Genus Stizoptera Oberholser, 1899
Stizoptera bichenovii (Vigors & Horsfield, 1827)

Genus Taeniopygia Reichenbach, 1862
Taeniopygia castanotis (Gould, 1837)
Taeniopygia guttata (Vieillot, 1817)

Genus Zonaeginthus Cabanis, 1851
Zonaeginthus bellus (Latham, 1801)
Zonaeginthus oculatus (Quoy & Gaimard, 1830)

LeNomenclatoriste said:

My new classification of Estrildidae based on their result

Click to expand...

The proposed classification in the paper seems rather inconsistent.

If we are to continue recognising (e.g.) Euschistospiza, Nesocharis, Padda and to elevate (e.g.) Glaucestrilda, Brunhilda, Mayrimunia... then lineages such as (e.g.) Rhodopyga, Granatina and Zonogastris, which are significantly older than any of these, should also be recognised at the generic level.

I'd prefer fewer genera overall as this is a fairly recent radiation - the whole group is less than 10.5 million years old.

For IOC to make the least changes:

- recognize Aidemosyne modesta, Bathilda ruficauda, Stizoptera bichenovii as species in monotypic genuses
- recognize Spermestes (Weird that they haven't yet)

- merge Nesocharis capistrata into Estrilda
OR
- split Estrilda in 3, given the age of the other genuses in Estrildinae: Delacourella, Brunhilda (incl. Glaucestrilda) & Estrilda

I think it makes sense to merge the remaining Nesocharis into Coccopygia

TomDerutter said:

For IOC to make the least changes

Click to expand...

I think that would preferable to adopting all the recommendations of the paper.

Using Olsson & Alström's tree, and applying a (conservative) 6my age requirement for a genus, you get:

Amandavi-
Ortygospiza
Paludipasser (presumably)
Amadina
Amandava (incl. Sporaeginthus)
Stictospiza

Lagonosticti-
Pytilia (incl. Zonogastris)
Lagonosticta (incl. Mormolycia & Rhodopyga)
Clytospiza
Hypargos (incl. Euschistospiza & Didymosticta)
Pyrenestes (incl. Spermophaga) (I'm not sure of priority here)
Uraeginthus (incl. Granatina)

Estrildi-
Cryptospiza (incl. Mandingoa)
Coccopygia (incl. Nesocharis)
Nigrita (incl. Parmoptila)
Estrilda (incl. Delacourella, Glaucestrilda, Brunhilda & Krimhilda)

Erythruri-
Erythrura (presumably incl. Rhamphostruthus)
Chloebia

Poephili-
Oreostruthus
Stagonopleura (incl. Zonaeginthus)
Neochmia (incl. Aegintha)
Emblema (incl. Bathilda & Aidemosyne)
Poephila (incl. Stizoptera & Taeniopygia)
Heteromunia

Lonchuri-
Spermestes
Odontospiza
Lepidopygia
Euodice
Lonchura (incl. Mayrimunia & Padda)

And no genus is as heterogeneous as the similarly-aged Vidua...

LeNomenclatoriste said:

My new classification of Estrildidae based on their result:

Family Estrildidae Bonaparte, 1850
Subfamily Estrildinae Bonaparte, 1850

Genus Brunhilda Reichenbach, 1862
Brunhilda charmosyna (Reichenow, 1881)
Brunhilda erythronotos (Vieillot, 1817)

Genus Coccopygia Reichenbach, 1863
Coccopygia bocagei Shelley, 1903
Coccopygia melanotis (Temminck, 1823)
Coccopygia quartinia (Bonaparte, 1850)

Genus Cryptospiza Salvadori, 1884
Cryptospiza jacksoni Sharpe, 1902
Cryptospiza reichenowii (Hartlaub, 1874)
Cryptospiza salvadorii Reichenow, 1892
Cryptospiza shelleyi Sharpe, 1902

Genus Delacourella Wolters, 1949
Delacourella capistrata (Hartlaub, 1861)

Genus Estrilda Swainson, 1827
Estrilda astrild (Linnaeus, 1758)
Estrilda atricapilla Verreaux & Verreaux, 1851
Estrilda kandti Reichenow, 1902
Estrilda melpoda (Vieillot, 1817)
Estrilda nigriloris Chapin, 1928
Estrilda nonnula Hartlaub, 1883
Estrilda ochrogaster Salvadori, 1897
Estrilda paludicola von Heuglin, 1863
Estrilda poliopareia Reichenow, 1902
Estrilda rhodopyga Sundevall, 1850
Estrilda rufibarba (Cabanis, 1851)
Estrilda troglodytes (Lichtenstein, 1823)

Genus Glaucestrilda Roberts, 1922
Glaucestrilda caerulescens (Vieillot, 1871)
Glaucestrilda perreini (Vieillot, 1817)
Glaucestrilda thomensis (de Sousa, 1888)

Genus Mandingoa Hartert, 1919
Mandingoa nitidula (Hartlaub, 1865)

Genus Nesocharis Alexander, 1903
Nesocharis ansorgei (Hartert, 1899)
Nesocharis shelleyi Alexander, 1903

Genus Nigrita Strickland, 1843
Nigrita bicolor (Hartlaub, 1844)
Nigrita canicapillus (Strickland, 1841)
Nigrita fusconotus Fraser, 1843
Nigrita luteifrons Verreaux & Verreaux, 1851

Genus Parmoptila Cassin, 1859
Parmoptila jamesoni (Shelley, 1890)
Parmoptila rubrifrons (Sharpe & Ussher, 1872)
Parmoptila woodhousei Cassin, 1859

Subfamily Lagonostictinae Steiner, 1960

Genus Clytospiza Shelley, 1896
Clytospiza monteiri (Hartlaub, 1860)

Genus Euschistospiza Wolters, 1943
Euschistospiza cinereovinacea (de Sousa, 1889)
Euschistospiza dybowskii (Oustalet, 1892)

Genus Granatina Sharpe, 1890
Granatina granatina (Linnaeus, 1766)
Granatina ianthinogaster (Reichenow, 1879)

Genus Hypargos Reichenbach, 1862
Hypargos margaritatus (Strickland, 1844)
Hypargos niveoguttatus (Peters, 1868)

Genus Lagonosticta Cabanis, 1851
– Subgenus Hypargoides Wolters, 1943
Lagonosticta (Hypargoides) nitidula Hartlaub, 1886
Lagonosticta (Hypargoides) rufopicta (Fraser, 1843)
Lagonosticta (Hypargoides) senegala (Linnaeus, 1766)
– Subgenus Lagonosticta Cabanis, 1851
Lagonosticta (Lagonosticta) landanae Sharpe, 1890
Lagonosticta (Lagonosticta) rhodopareia (von Heuglin, 1868)
Lagonosticta (Lagonosticta) rubricata (Lichtenstein, 1823)
Lagonosticta (Lagonosticta) sanguinodorsalis Payne, 1998
Lagonosticta (Lagonosticta) umbrinodorsalis Reichenow, 1910
Lagonosticta (Lagonosticta) virata Bates, 1932
– Subgenus Lychnidopsiza Heuglin, 1871
Lagonosticta (Lychnidospiza) rara (Antinori, 1864)
– Subgenus Mormolycia Reichenbach, 1862
Lagonosticta (Mormolycia) larvata (Rüppell, 1840)
Lagonosticta (Mormolycia) nigricollis Heuglin, 1863
Lagonosticta (Mormolycia) vinacea (Hartlaub, 1857)

Genus Pyrenestes Swainson, 1837
Pyrenestes minor Shelley, 1894
Pyrenestes ostrinus (Vieillot, 1805)
Pyrenestes sanguineus Swainson, 1837

Genus Pytilia Swainson, 1837
Pytilia afra (Gmelin, 1789)
Pytilia hypogrammica Sharpe, 1870
Pytilia lineata von Heuglin, 1863
Pytilia phoenicoptera Swainson, 1837

Genus Spermophaga Swainson, 1837
Spermophaga haematina (Vieillot, 1808)
Spermophaga poliogenys (Ogilvie-Grant, 1906)
Spermophaga ruficapilla (Shelley, 1888)

Genus Uraeginthus Cabanis, 1851
Uraeginthus angolensis (Linnaeus, 1758)
Uraeginthus bengalus (Linnaeus, 1766)
Uraeginthus cyanocephalus (Richmond, 1897)

Genus Zonogastris Cabanis, 1851
Zonogastris melba (Linnaeus, 1758)

Sous-famille Amandavinae Steiner, 1960

Genus Amadina Swainson, 1827
Amadina erythrocephala (Linnaeus, 1758)
Amadina fasciata (Gmelin, 1789)

Genus Amandava Blyth, 1836
Amandava amandava (Linnaeus, 1758)

Genus Ortygospiza Sundevall, 1850
Ortygospiza atricollis (Vieillot, 1817)
Ortygospiza fuscocrissa von Heuglin, 1863
Ortygospiza gabonensis Lynes, 1914

Genus Paludipasser Neave, 1909
Paludipasser locustella Neave, 1909

Genus Sporaeginthus Cabanis, 1851
Sporaeginthus subflavus (Vieillot, 1819)

Genus Stictospiza Sharpe, 1890
Stictospiza formosa (Lathamn, 1790)

Subfamily Erythrurinae Delacour, 1943

Genus Chloebia Reichenbach, 1862
Chloebia gouldiae (Gould, 1844)

Genus Erythrura Swainson, 1837
– Subgenus Amblynura Reichenbach, 1862
Erythrura (Amblynura) coloria Ripley & Rabor, 1961
Erythrura (Amblynura) cyanovirens (Peale, 1848)
Erythrura (Amblynura) papuana Rothschild & Hartert, 1900
Erythrura (Amblynura) pealii Hartlaub, 1852
Erythrura (Amblynura) psittacea (Gmelin, 1789)
Erythrura (Amblynura) regia (Sclater, 1881)
Erythrura (Amblynura) tricolor (Vieillot, 1817)
Erythrura (Amblynura) trichroa (von Kittlitz, 1833)
– Subgenus Erythrura Swainson, 1837
Erythrura (Erythrura) prasina (Sparrman, 1788)
Erythrura (Erythrura) viridifacies Hachisuka & Delacour, 1937
– Subgenus Reichenowia Poche, 1904
Erythrura (Reichenowia) hyperythra (Reichenbach, 1862)
– Subgenus Rhamphostruthus Mayr, 1931
Erythrura (Rhamphostruthus) kleinschmidti (Finsch, 1878)

Subfamily Lonchurinae Steiner, 1960

Genus Euodice Reichenbach, 1862
Euodice cantans (Gmelin, 1789)
Euodice malabarica (Linnaeus, 1758)

Genus Lepidopygia Reichenbach, 1862
Lepidopygia nana (Pucheran, 1845)

Genus Lonchura Sykes, 1832
– Subgenus Lonchura Sykes, 1832
Lonchura (Lonchura) punctulata (Linnaeus, 1758)
– Subgenus Munia Hodgson, 1836
Lonchura (Munia) atricapilla (Vieillot, 1807)
Lonchura (Munia) caniceps (Salvadori, 1876)
Lonchura (Munia) castaneothorax (Gould, 1837)
Lonchura (Munia) ferruginosa (Sparrman, 1789)
Lonchura (Munia) flaviprymna (Gould, 1845)
Lonchura (Munia) forbesi (Sclater, 1879)
Lonchura (Munia) grandis (Sharpe, 1882)
Lonchura (Munia) hunsteini (Finsch, 1886)
Lonchura (Munia) maja (Linnaeus, 1766)
Lonchura (Munia) malacca (Linnaeus, 1766)
Lonchura (Munia) melaena (Sclater, 1880)
Lonchura (Munia) montana Junge, 1939
Lonchura (Munia) monticola (De Vis, 1897)
Lonchura (Munia) nevermanni Stresemann, 1934
Lonchura (Munia) pallida (Wallace, 1864)
Lonchura (Munia) quinticolor (Vieillot, 1807)
Lonchura (Munia) spectabilis (Sclater, 1878)
Lonchura (Munia) stygia Stresemann, 1934
Lonchura (Munia) teerinki Rand, 1940
Lonchura (Munia) vana (Hartert, 1930)
– Subgenus Trichogrammoptila Reichenbach, 1862
Lonchura (Trichogrammoptila) fuscans (Cassin, 1852)
Lonchura (Trichogrammoptila) kelaarti (Jerdon, 1863)
Lonchura (Trichogrammoptila) leucogastra (Blyth, 1846)
Lonchura (Trichogrammoptila) leucogastroides (Moore, 1858)
Lonchura (Trichogrammoptila) molucca (Linnaeus, 1758)
Lonchura (Trichogrammoptila) striata (Linnaeus, 1766)

Genus Mayrimunia Wolters, 1949
Mayrimunia leucostista (d'Albertis & Salvadori, 1879)
Mayrimunia tristissima (Wallace, 1865)

Genus Odontospiza Oberholser, 1905
Odontospiza griseicapilla (Delacour, 1943)

Genus Padda Bonaparte, 1850
Padda fuscata (Vieillot, 1807)
Padda oryzivora (Linnaeus, 1758)

Genus Spermestes Swainson, 1837
Spermestes bicolor (Fraser, 1843)
Spermestes cucullata Swainson, 1837
Spermestes fringilloides (Lafresnaye, 1835)

Subfamily Poephilinae Mayr, Paynter & Taylor, 1968

Genus Aegintha Cabanis, 1851
Aegintha temporalis (Latham, 1801)

Genus Aidemosyne Reichenbach, 1862
Aidemosyne modesta (Gould, 1837)

Genus Bathilda Reichenbach, 1862
Bathilda ruficauda (Gould, 1837)

Genus Emblema Gould, 1842
Emblema pictum Gould, 1842

Genus Heteromunia Mathews, 1913
Heteromunia pectoralis (Gould, 1841)

Genus Neochmia Gray, 1849
Neochmia evangelinae d'Albertis & Salvadori, 1879
Neochmia phaeton (Hombron & Jacquinot, 1841)

Genus Oreostruthus De Vis, 1898
Oreostruthus fuliginosus (De Vis, 1897)

Genus Poephila Gould, 1842
Poephila acuticauda (Gould, 1840)
Poephila cincta (Gould, 1837)
Poephila personata Gould, 1842

Genus Stagonopleura Reichenbach, 1850
Stagonopleura guttata (Shaw, 1796)

Genus Stizoptera Oberholser, 1899
Stizoptera bichenovii (Vigors & Horsfield, 1827)

Genus Taeniopygia Reichenbach, 1862
Taeniopygia castanotis (Gould, 1837)
Taeniopygia guttata (Vieillot, 1817)

Genus Zonaeginthus Cabanis, 1851
Zonaeginthus bellus (Latham, 1801)
Zonaeginthus oculatus (Quoy & Gaimard, 1830)

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Rubropyga, of course, and not Hypargoides

I'm quite extremist by recognising Krimhilda (kandti, atricapilla and nonnula) as distinct genus from Estrilda.

Didymosticta ? never heard until now

Poephili-
Oreostruthus
Stagonopleura (incl. Zonaeginthus)
Neochmia (incl. Aegintha)
Emblema (incl. Bathilda & Aidemosyne)
Poephila (incl. Stizoptera & Taeniopygia)

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This seems better then having 7 (out of 10) monotypic genera

Still need Heteromunia there

Could go a little further to 7mya. (You start losing meaningfull info on this level probably)

Amandavi-
Ortygospiza
Paludipasser (presumably)
Amadina
Amandava (incl. Sporaeginthus & Stictospiza)

Lagonosticti-
Pytilia (incl. Zonogastris)
Lagonosticta (incl. Mormolycia & Rhodopyga)
Clytospiza
Hypargos (incl. Euschistospiza & Didymosticta)
Pyrenestes (or Spermophaga - priority?) (incl. Spermophaga, Uraeginthus, Granatina)

Estrildi-
Cryptospiza (incl. Mandingoa)
Coccopygia (incl. Nesocharis)
Nigrita (incl. Parmoptila)
Estrilda (incl. Delacourella, Glaucestrilda, Brunhilda & Krimhilda)

Erythruri-
Erythrura (incl. Chloebia & presumably Rhamphostruthus)

Poephili-
Heteromunia
Oreostruthus
Stagonopleura (incl. Zonaeginthus)
Emblema (or Poephila - priority?) (incl. Neochmia, Aegintha, Bathilda & Aidemosyne, Poephila, Stizoptera & Taeniopygia)

Lonchuri-
Spermestes (incl. Odontospiza)
Lonchura (incl. Lepidopygia, Euodice, Mayrimunia & Padda)

TomDerutter said:

This seems better then having 7 (out of 10) monotypic genera

Still need Heteromunia there

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Oh yeah - I forgot Heteromunia - I'll edit

A couple of family-group name issues...

• Estrildinae (Bonaparte, 1850) (clade A). Referred taxa: Brunhilda, Coccopygia, Cryptospiza, Estrilda, Delacourella, Glaucestrilda, Mandingoa, Nesocharis, Nigrita and Parmoptila.

In my notes, Estrildidae/Estrildinae dates from Blyth 1846, not from Bonaparte 1850.
- Estrildinae (subfamily) : Blyth E. 1846. Notices and descriptions of various new or little known species of birds. [Continued from Vol. XIV, p.602]. J. Asiat. Soc. Bengal 15: 1-54.; p. 36; https://biodiversitylibrary.org/page/40122571

• Lagonostictinae (Steiner, 1960) (clade B). Referred taxa: Clytospiza, Euschistospiza, Granatina, Hypargos, Lagonosticta, Pyrenestes, Pytilia, Spermophaga and Uraeginthus.

Steiner 1960 used an apparent plural “Lagonostictae” only once in his text, when discussing his so-called "Tribus Didymostictae", in a sentence saying that the species [Clytospiza] cinereo-vinacea of this group leads to "Lagonostictae". He did not adopt such a group in his classification at a supra-generic rank; he placed Lagonosticta in Tribus Estrildae, which he directly subdivided into two genera, Lagonosticta and Estrilda; under Estrildae, Steiner re-affirmed a link beween Clyptospiza cinereo-vinacea and "Lagonosticta", this time using the generic name in the singular. I do not see that "Lagonostictae" was "clearly used as a scientific name to denote a suprageneric taxon and not merely as a plural noun or adjective referring to the members of a genus" (11.7.1.2.) by Steiner and, if not, it is not a family-group name. Additionally, the purported taxon was not given a diagnosis, in a work published after 1830, and I am unaware that the name was adopted by any subsequent author before 2000 : thus I'm not at all sure that the exception offered by Art 13.2.1 for family-group names published after 1830 but before 1961 can be deemed to apply. My current impression is that this name doesn't exist at all.
Why not Pyrenestinae Bonaparte 1854 ? (Please not "Pytiliinae Bonaparte 1840".)
- "Lagonostictae" (not adopted at any suprageneric rank) : Steiner H. 1960. Klassifikation der Prachtfinken, Spermestidae, auf Grund der Rachenzeichnungen ihrer Nestlinge. J. Ornithol. 101: 92-112; p. 96; https://doi.org/10.1007/BF01670634
- Pyrenesteae (group) : Bonaparte CL. 1854. Conspectus systematis ornithologiae. Ann. Sci. Nat. Zool., sér.4, 1: 105-152.; p. 115; http://biodiversitylibrary.org/page/35982400
- ("Pytiliidae Bonaparte 1840" apud Bock 1994 http://digitallibrary.amnh.org/handle/2246/830 = Pytilinae (subfamily) : Bonaparte CL. 1840. A new systematic arrangement of the vertebrate animals. Trans. Linn. Soc. London 18: 247-304.; p. 271; http://biodiversitylibrary.org/page/4862456 ; see the diagnosis: short, thick, subtriangular bill, 9 primaries only... The type of this group is obviously not the delicate and 10-primaried Estrildid Pytilia Swainson, it is Pitylus Cuvier 1829. I have yet to see an available family-group name based on Pytilia.)

• Amandavinae (Steiner, 1960) (clade C). Referred taxa: Amadina, Amandava, Ortygospiza and tentatively Paludipasser.

This one seems available from Steiner 1960, but it is not currently in use. Why should the senior name Amadininae Bonaparte 1854 be discarded ?
- Amandavae (tribe) : Steiner H. 1960. Klassifikation der Prachtfinken, Spermestidae, auf Grund der Rachenzeichnungen ihrer Nestlinge. J. Ornithol. 101: 92-112; p. 103; https://doi.org/10.1007/BF01670634
- Amadineae (group) : Bonaparte CL. 1854. Conspectus systematis ornithologiae. Ann. Sci. Nat. Zool., sér.4, 1: 105-152.; p. 115; http://biodiversitylibrary.org/page/35982400

• Erythrurinae (Delacour, 1943) (clade D). Referred taxa: Chloebia and Erythrura.

This one is OK.
- Erythrurae (tribe) : Delacour J. 1943. A revision of the subfamily Estrildinae of the family Ploceidae. Zoologica (New York) 28: 69-86.; p. 71; http://biodiversitylibrary.org/page/51008604

• Lonchurinae (Steiner, 1960) (clade E). Referred taxa: Euodice, Lepidopygia, Lonchura, Mayrimunia, Padda, and Spermestes.

This name is available from Steiner 1960 too, and is in use, so I guess it would be best retained... I'm not that sure this is what the Code says, though. (Bock 1994 treated it as taking precedence from its senior synonym Spermestinae Cabanis 1847 as per what is now Art. 40.2 (http://digitallibrary.amnh.org/handle/2246/830; p. 156 : "LONCHURINAE Steiner, 1960 (1847)"); however, this would be true only if Cabanis' name had been "replaced before 1961 because of the synonymy of the type genus"; Steiner 1960 (see the work's title) created a new tribe Lonchurae which he placed in the family Spermestidae, thus he did not "replace" this last name with the first one in any meaningful way. All the subsequent uses of the name date from after 1960, and are thus too late for 40.2 to apply. Spermestidae can certainly not be claimed to be a nomen oblitum.)
- Lonchurae (tribe) : Steiner H. 1960. Klassifikation der Prachtfinken, Spermestidae, auf Grund der Rachenzeichnungen ihrer Nestlinge. J. Ornithol. 101: 92-112; p. 96, 107; https://doi.org/10.1007/BF01670634
- Spermestinae (subfamily) : Cabanis J. 1847. Ornithologische Notizen. Arch. Naturgesch. 13: 186-256;308-352.; p. 331; http://biodiversitylibrary.org/page/14785301

• Poephilinae (Mayr, Paynter & Traylor, 1968) (clade F). Referred taxa: Aidemosyne, Bathilda, Emblema, Heteromunia, Neochmia, Oreostruthus, Poephila, Stagonopleura, Stizoptera and Taeniopygia.

Introduced after 1960 without a single word of description -- the name, as used by Mayr, Paynter & Traylor 1968, is a pure nomen nudum, it can certainly not be cited from there. Is there a better source (with a decent diagnonis) somewhere else ?
Zonaeginthinae Steiner 1960 applies to the same group (Zonaeginthus is a syn. of Stagonopleura in the adopted classification), and is available.
- Poephilae (unranked group) : Mayr E, Paynter RA, Traylor MA. 1968. Family Estrildidae. Pp. 306-390 in: Paynter RA (Ed.). 1968. Check-list of birds of the world. A continuation of the work of James L. Peters. Volume XIV. Mus. Comp. Zool., Cambridge, MA.; p. 352; http://biodiversitylibrary.org/page/14481553
- Zonaeginthae (tribe) : Steiner H. 1960. Klassifikation der Prachtfinken, Spermestidae, auf Grund der Rachenzeichnungen ihrer Nestlinge. J. Ornithol. 101: 92-112; p. 106; https://doi.org/10.1007/BF01670634

LeNomenclatoriste said:

Didymosticta ? never heard until now

Click to expand...

What is Rubropyga?

l_raty said:

What is Rubropyga?

Click to expand...

A typo. I mean Rhodopyga

TomDerutter said:

Pyrenestes (or Spermophaga - priority?)

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Both names have been quoted from the same work: Swainson W. 1837. Birds of Western Africa. [1st part.] Ornithology. Vol. VII. In: Jardine W [ed]. The naturalist's library. WH Lizars, Edinburgh. Pyrenestes : p. 156; https://books.google.com/books?id=9I0-AAAAcAAJ&pg=PA156 (in the header spelled Pirenestes; in the text Pyrenestes); Spermophaga : p. 164; https://books.google.com/books?id=9I0-AAAAcAAJ&pg=PA164. If this is so, the date of publication is the same, and precedence is not fixed originally -- you need a first reviser act, which may not actually exist.
I'm inclined to take Pyrenestes from an earlier work, however: Swainson W. 1836. On the natural history and classification of birds. Vol. I. In: Lardner D [ed]. The cabinet cyclopaedia. Longman, Rees, Orme, Brown, Green, & Longman, London.; p. viii; https://biodiversitylibrary.org/page/46085808 (associated to a woodcut on p. 65 of the work https://biodiversitylibrary.org/page/42333403). If taken from there, it has clear precedence.

Emblema (or Poephila - priority?)

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Poephila first, in Part 6 of Gould's Birds of Australia, 1 Mar 1842; then Emblema, in Part 7, 1 Jun 1842. (Cf. Waterhouse 1885 https://biodiversitylibrary.org/page/12782774, https://biodiversitylibrary.org/page/12782744; Zimmer 1926 https://biodiversitylibrary.org/page/36194906. Both names also in PZS, in Nov of the same year.)

DECICCO, L. H., BENZ, B. W., DERAAD, D. A., HIME, P. M. & MOYLE, R. G. New Guinea Erythrura parrotfnches: one species or two? Bull. B.O.C. 2020 140(3).

Summary.—Two species of Erythrura parrotfnches, difering mainly in bill size,
are described from the New Guinea highlands: Blue-faced Parrotfnch E. trichroa
and Papuan Parrotfnch E. papuana. Morphological measurements from museum
specimens support two non-overlapping groups, but mitochondrial DNA sequence
data show negligible diferences between the two species. These observations
suggest that E. trichroa and E. papuana may form a single species in the highlands
of New Guinea that exhibits a resource-based bill size polymorphism.

Lopez, K.A., C.S. McDiarmid, S.C. Griffith, I.J. Lovette, and D.M. Hooper (2021)
Evaluating evidence of mitonuclear incompatibilities with the sex chromosomes in an avian hybrid zone
Evolution (advance online publication)
doi: 10.1111/evo.14243
https://onlinelibrary.wiley.com/doi/10.1111/evo.14243

The exploration of hybrid zones and the intergenomic conflicts exposed through hybridization provide windows into the processes of divergence and speciation. Sex chromosomes and mitonuclear incompatibilities have strong associations with the genetics of hybrid dysfunction. In ZW sex‐determining systems, maternal co‐inheritance of the mitochondrial and W chromosomes immediately exposes incompatibilities between these maternal contributions of one species and the Z chromosome of another. We analyze mitochondrial and Z chromosome admixture in the long‐tailed finch (Poephila acuticauda) of Australia, where hybridizing subspecies differ prominently in Z chromosome genotype and in bill color, yet the respective centers of geographic admixture for these two traits are offset by 350 km. We report two well‐defined mitochondrial clades that diverged ∼0.5 Mya. Mitochondrial contact is geographically co‐located within a hybrid zone of Z chromosome admixture and is displaced from bill color admixture by nearly 400 km. Consistent with Haldane's rule expectations, hybrid zone females are significantly less likely than males to carry an admixed Z chromosome or have mismatched Z‐mitochondrial genotypes. Furthermore, there are significantly fewer than expected mitonuclear mismatches in hybrid zone females and paternal backcross males. Results suggest a potential for mitonuclear/sex chromosome incompatibilities in the emergence of reproductive isolation in this system.

Rivera, Moises, Edwards, Jacob A., Hauber, Mark E. and Sarah M. N. Wooley (2023) Machine learning and statistical classification of birdsong link vocal acoustic features with phylogeny. Scientific Reports 13: 7076. Machine learning and statistical classification of birdsong link vocal acoustic features with phylogeny - Scientific Reports

Abstract
Birdsong is a longstanding model system for studying evolution and biodiversity. Here, we collected and analyzed high quality song recordings from seven species in the family Estrildidae. We measured the acoustic features of syllables and then used dimensionality reduction and machine learning classifiers to identify features that accurately assigned syllables to species. Species differences were captured by the first 3 principal components, corresponding to basic frequency, power distribution, and spectrotemporal features. We then identified the measured features underlying classification accuracy. We found that fundamental frequency, mean frequency, spectral flatness, and syllable duration were the most informative features for species identification. Next, we tested whether specific acoustic features of species’ songs predicted phylogenetic distance. We found significant phylogenetic signal in syllable frequency features, but not in power distribution or spectrotemporal features. Results suggest that frequency features are more constrained by species’ genetics than are other features, and are the best signal features for identifying species from song recordings. The absence of phylogenetic signal in power distribution and spectrotemporal features suggests that these song features are labile, reflecting learning processes and individual recognition.

Lucas H. DeCicco, Devon A. DeRaad, Emily N. Ostrow, Robert G. Moyle (2023). A complete species-level phylogeny of the Erythrura parrotfinches (Aves: Estrildidae), Molecular Phylogenetics and Evolution, 107883.


Abstract: Identifying species boundaries and phylogenetic relationships among groups of closely related species provides a necessary framework for understanding how biodiversity evolves in natural systems. Here we present a complete phylogeny of the avian genus Erythrura (family Estrildidae) commonly known as parrotfinches, which includes species threatened by habitat loss and the pet trade. Using both mitogenome and reduced-representation genome-wide nuclear DNA sequence data, we reconstructed the evolutionary history of the group by sampling all 12 recognized species, four of which had not previously been studied in a phylogenetic context. We included intra-species geographic sampling that allowed us to comment on species limits in some taxa. We recovered the Gouldian Finch (Chloebia gouldiae) of Australia which has often been placed in the monotypic genus Chloebia, as being sister to a clade comprising all Erythrura species. In addition, we recovered a well-supported clade comprising eight species distributed throughout the Pacific Island eco-region, whereas those species occurring in continental southeast Asian, the Greater Sundas, and the Philippines, were recovered as earlier branching lineages. Of note was the early branching of the Fiji-endemic E. kleinschmidti which corroborates its unique phenotype. We also found a deep phylogenetic split (8.59% corrected, 7.89% uncorrected divergence in the mitochondrial gene ND2) between the Java and Philippine populations of E. hyperythra, indicating unrecognized species-level diversity within this taxon. In contrast, genome-wide nuclear data suggested that the New Guinea endemic species E. papuana is embedded within the widespread species E. trichroa in all phylogenetic reconstructions, corroborating previously published mitochondrial data that suggested a similar pattern. By generating a phylogenetic hypothesis for the relationships among all species of Erythrura parrotfinches, we provide a framework for better understanding the extant diversity and evolutionary history of this group.

I keep my current taxonomy, e.g. Chloebia, Erythrura and Rhamphostruthus

Question on type species of Lonchura

Background

The genus Lonchura (family Estrildidae) was introduced by Sykes in 1832 here. Sykes listed three species in the genus:

• Lonchura nisoria, Fringilla nisoria, Temm
• Lonchura cheet
• Lonchura leuconota, Fringilla leuconota, Temm

Fringilla nisoria had been described by Temminck in 1930 here

Fringilla nisoria is now considered as a subspecies of the Scaly-breasted Munia (Lonchura punctulata nisoria)

In 1890 Sharpe fixed the type of Lonchura as Munia punctulata here:

H&M4 Vol 2 p. 297 available here includes the text:

"LONCHURA Sykes, 1832 F - Fringilla nisoria Sykes, 1832; type by subsequent designation (Sharpe, 1890, Cat. Birds Brit. Mus., 13, p. 326). = Loxia punctulata Linnaeus, 1758"

Questions

1. Why does H&M4 give the authority for Fringilla nisoria as Sykes 1832 rather than Temminck 1830? Is this an error?

2. Is the type equivalent to the subspecies Lonchura punctulata nisoria or to the nominate Lonchura punctulata punctulat?.

Peter Brick said:

Fringilla nisoria had been described by Temminck in 1930 here

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1830 x)