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Greater White-fronted Geese (1 Viewer)

Papuan birder

- Lost in the Pacific -
Taxonomy of Greater White-fronted Geese (Aves: Anatidae)

Five subspecies of the Greater White-fronted Goose, Anser albifrons (Scopoli, 1769), have been named, all on the basis of wintering birds, and up to six subspecies have been recognized. There has been confusion over the application of some names, particularly in North America, because of lack of knowledge of the breeding ranges and type localities, and incorrect taxonomic decisions. There is one clinally varying subspecies in Eurasia, one that breeds in Greenland, and three in North America, one newly named herein.

http://www.bioone.org/doi/abs/10.2988/11-14.1
 
Anser albifrons

Taxonomy of Greater White-fronted Geese (Aves: Anatidae)

Five subspecies of the Greater White-fronted Goose, Anser albifrons (Scopoli, 1769), have been named, all on the basis of wintering birds, and up to six subspecies have been recognized. There has been confusion over the application of some names, particularly in North America, because of lack of knowledge of the breeding ranges and type localities, and incorrect taxonomic decisions. There is one clinally varying subspecies in Eurasia, one that breeds in Greenland, and three in North America, one newly named herein.

http://www.bioone.org/doi/abs/10.2988/11-14.1

Systematics:
1. Anser albifrons albifrons (Scopoli, 1769)
Distribution: Breeds in northern Eurasia
2. Anser albifrons flavirostris Dalgety & Scott, 1948
Distribution: Breeds in western Greenland
3. Anser albifrons elgasi Delacour & Ripley, 1975
Distribution: Breeds in the Cook Inlet area of southern Alaska
4. Anser albifrons gambelli Hartlaub, 1852
Distribution: Breeds in interior and northern Alaska and across arctic Canada to the Hudson Bay region
5. Anser albifrons sponsa, Banks, 2011 (new ssp)
Distribution: Breeds in western Alaska, in the Yukon-Kuskokwim Delta area and in the Bristol Bay lowlands
 
So, frontalis Baird, 1858 not recognised; and gambelli rather than gambeli (eg, Mayr & Cottrell 1979, H&M3).

Haven't seen the paper, but why isn't sponsa a junior synonym of frontalis (described from NM, but supposedly breeding w/n AK & e Siberia)?

See also this older thread:
www.birdforum.net/showthread.php?t=152366
 
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and gambelli rather than gambeli (eg, Mayr & Cottrell 1979, H&M3).
Alan Peterson explains it well. I may add that this is a good example of why "prevailing usage" is a silly idea. Google Scholar gives 466 hits for gambeli, but also 127 hits for gambelli, many of which are from the last 20 years. There's a majority for gambeli, but who may judge if it's prevailing? And how?

If I were asked I'd recommend use of the original spelling gambelli.

Rainer
 
Frontalis

Haven't seen the paper, but why isn't sponsa a junior synonym of frontalis (described from NM, but supposedly breeding w/n AK & e Siberia)?
Suggests that the NM individual was probably a stray from a population migrating to winter in Texas, and frontalis must be considered a synonym of gambelli. (A pitfall of describing migrant taxa away from the breeding range.)
 
'Albicans'

Further to my earlier question about albicans...
The population of eastern Eurasia was distinguished from the western nominate form by Mooij (2000:386) and Mooij & Zöckler (2000:101) on the basis of its somewhat larger size. Those authors used the subspecific name A. a. albicans "according to Gmelin 1788, quoted by Alphéraky (1904)." Gmelin (1788:416) listed the name albicans only in reference to "Branta albifrons Scop.," of which it is clearly a synonym, and gave no indication, at any rate, that it would apply to birds of eastern Asia. The name is mentioned by Alphéraky (1905:42) only in the extensive synonymy of the species Anser albifrons as "Anas albicans (errore), Gmelin" which is as it was cited by Salvadori (1895:92). It is clearly not an available name (ICZN 1999) and cannot be applied to any population of Anser albifrons. Delacour (1954), and those who followed him, also divided the Eurasian population into two, but with different boundaries, referring the eastern birds to the North American A. a. frontalis.
 
Pacific White-fronted Geese

Some interesting ID notes on Martin Garner's Birding Frontiers blog: Pacific White-fronted Goose?

Still tentatively using the names albicans (which Banks 2011 has confirmed to be incorrect) and frontalis (synonymised with gambelli by Banks) for Asian and American populations respectively. Banks assigns all Eurasian birds to nominate albifrons - presumably a new name is needed if East Asian birds are to be treated as a distinct ssp.
 
Suggests that the NM individual was probably a stray from a population migrating to winter in Texas, and frontalis must be considered a synonym of gambelli. (A pitfall of describing migrant taxa away from the breeding range.)

"Baird 1858 frontalis is from a young bird from Fort Thorn New Mexico. Possible a wintering siberian bird, but in 1859 Baird added a bird from the Selkirk Settlement Manitoba as a co-type? A breeding bird from the plains of Canada not siberia. "
Banks mentions the bird from the Selkirk Settlement Manitoba but does not address it by saying Cassin listed the Fort Thorn New Mexico bird as the type (Art. 74.5) Is it as likely the New Mexico bird was from some Pacific population as a Texas bird?

74.5. Lectotype designations before 2000. In a lectotype designation made before 2000, either the term "lectotype", or an exact translation or equivalent expression (e.g. "the type"), must have been used or the author must have unambiguously selected a particular syntype to act as the unique name-bearing type of the taxon. When the original work reveals that the taxon had been based on more than one specimen, a subsequent use of the term "holotype" does not constitute a valid lectotype designation unless the author, when wrongly using that term, explicitly indicated that he or she was selecting from the type series that particular specimen to serve as the name-bearing type.
 
gambelli rather than gambeli .


Banks says “The International Code of Zoological Nomenclature (ICZN
1999), Article 32, states that the original spelling of a name used in the work in
which the name was established is the ‘‘correct original spelling’’ and must be
used unless there is, in the original publication itself, clear evidence of an
inadvertent error. As McAtee (1944) VOLUME 124, NUMBER 3 231pointed out there is no internal evidence in Hartlaub’s (1852) paper to support Coues’s (1882) assertion, (that the goose was named for William Gambel) But there is evidence contemporaneous in time, place and publication that it was in fact named for William Gambel.
“In Naumannia, Volume 2 (1852); was an article by August Karl Eduard Baldamus, Auszug aus dem Protokolle der Versammlung der deutschen Ornithologen zu Berlin 1851, which reported on an oral presentation by Hartlaub accompanied by an illustration! Wonder where that drawing is? (The article was dated June 1851 but probably published in 1852)

“Dr. Hartlaub of Bremen spoke about a new North American goose, which the European albifrons is very close and, it seems, Bonaparte and Audubon, with the same confused by Wilson's been. The same is different but most definitely by far stronger beak dimensions, as explained, a drawing by. Example three. who knows by Hartlaub, two are from Texas and one from southern your part of the United States. It was for this new goose the name Anser Gambelli proposed, in recognition of the merits of American zoologists Gambel to the ornithology of California.” .”
I do not interpret the code to require such information (internal evidence in Hartlaub’s (1852) paper ) in the original paper going towards the correctness of the initial spelling.


“If it is shown subsequently that the precedence of names, spellings or acts can be objectively determined, the action of the First Reviser is nullified. “

This article shows the code’s reasonableness. Even though we are not looking at a first reviser situation this shows only objective reasonableness is required and The second article is proof enough for the code that the bird was named for William Gamble.

32.5.1. If there is in the original publication itself, without recourse to any external source of information, clear evidence of an inadvertent error, such as a lapsus calami or a copyist's or printer's error, it must be corrected. Incorrect transliteration or latinization, or use of an inappropriate connecting vowel, are not to be considered inadvertent errors. This article does not apply to this situation. So I think the emendation of Cassin etc was correcting a misspelling. They worked together Gambel named the Cassin’s Auklet for Cassin!
 
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sponsa in Asia

In Banks 2011, is quoted:
“Ely et al. (2005) studied morphological variation (size) of Greater White-fronted Geese on their nesting grounds, and with consideration of other factors have provided a basis for dividing the species into subspecies. Briefly summarized, they found a clinal increase in size from west to east across Eurasia, a similarity of size of eastern Asian birds and those of the west coast of Alaska, without any indication of interchange of individuals between these areas, and little variation in samples from interior Alaska and across northern Canada.”

The italics are mine, as this interest me:
As the East Asian breeding birds are the same size as West Alaskan birds (the newly named ssp sponsa), they too must be sponsa (it having been described on the basis of size alone), surely?
 
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Banks 2012

Banks 2011 (Taxonomy of Greater White-fronted Geese (Aves: Anatidae)) pdf free download here.
Banks 2012. Geographic variation in wintering Greater White-fronted Geese. Western Birds 43(4).
ABSTRACT: There is relatively little variation in size, expressed mainly in bill dimensions, between or among most wintering populations of the Greater White-fronted Goose (Anser albifrons). In the British Isles, slightly larger and darker birds, from the Greenland breeding population, winter in Ireland and Scotland and associated islands, while smaller birds winter in England. Winter birds in continental Europe are the same size as those in England. Asian winter birds average slightly larger than those of Europe; the population is more variable and includes some larger individuals. In western North America, some birds in the Sacramento Valley of northern California, the famed Tule Goose (A. a. elgasi), are the largest of the species. There is a great range of variation in smaller birds of the Sacramento Valley and elsewhere in the west coast states. Birds in the midcontinent states, east of the Rocky Mountains, average about the same as smaller California birds but vary widely.
Presumably provides further justification for the recognition of sponsa Banks, 2011.
 
Eda et al 2013

Eda, Shimada, Amano, Ushiyama, Mizota & Koike 2013. Phylogenetic relationship of the Greater White-Fronted Goose Anser albifrons subspecies wintering in the Palaearctic region. Ornithol Sci 12(1): 35–42. [abstract]

Recognises frontalis, contra Banks 2011, OSJ 2012, H&M4, IOC.
 
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