Peter Kovalik
Well-known member
PASSED (6 June 2019)
Proposal (827) to SACC
Revise the classification and linear sequence of the Tyrannoidea: (A) Recognize a new family Onychorhynchidae; (B) Modify linear sequence of families; (C) Add three subfamilies to the Tyrannidae
Proposal (827) to SACC
Revise the classification and linear sequence of the Tyrannoidea: (A) Recognize a new family Onychorhynchidae; (B) Modify linear sequence of families; (C) Add three subfamilies to the Tyrannidae
Lavinia P.D., Escalante P., Tubaro P.L. & Lijtmaer D.A., 2020. Molecular phylogenetics and phenotypic reassessment of the Ramphotrigon flycatchers: deep paraphyly in the context of an intriguing biogeographic scenario. J. Avian Biol.
There
Ochthoecini, Ohlson, Irestedt, Batalha Filho, Ericson and Fjeldså, new family group name
Type genus: Ochthoeca Cabanis, 1847
Included genera: Ochthoeca Cabanis, Myiophobus Reichenbach, Colorhamphus Sundevall, Silvicultrix Lanyon, and Scotomyias, Ohlson, Irestedt, Batalho Filho, Ericson and Fjeldså.
Diagnosis: A morphologically homogeneous group, which is well-supported in all published phylogenetic studies. There are no known internal anatomical characters that define the clade. Morphologically and behaviorally the
group is well-defined as sexually monomorphic flycatchers with a compact body shape, generally upright posture with protruding breast when perched, and moderately long tails, moderately long tarsi, and a triangular, moderately compressed and broad-based bill with rictal bristles extending slightly beyond the middle of the bill. Plumage colouration and patterns generally simple. In general, colouration includes dark olive, earthy brownish or sooty upperparts and underparts varying from sooty grey to rufous, pale grey, dull yellowish or bright yellow depending on the species. Some species have a rufous breast band; streaking on the chest occurs only in Myiophobus. Myiophobus and Ochthoeca salvini have distinctive pale wing bars, otherwise wing patterns are usually subdued, mostly in the form of dull cinnamon or buff wing bars. Distinctive markings include bright white, yellow, or rufous bands across forehead and eyebrow (Ochthoeca, Silvicultrix) and bright yellow or orange coronal patches (Myiophobus, Scotomyias).
Cladistic definition: All descendants of the most recent common ancestor of Myiophobus fasciatus (Statius Müller) and Ochthoeca leucophrys (d’Orbigny & Lafresnaye).
Habitat and distribution: Myiophobus has a wide distribution in scrubby habitats in tropical lowlands of South America; the remainder of the genera are largely restricted to the Andes, with one species in the Tumbesian lowlands, one species in the tepuis and one species in the austral Nothofagus forest. Most species occur in humid forest undergrowth and edges, but one clade in Ochthoeca favours drier habitats. All species forage by short sallies to air or vegetation, usually from a low perch.
The name Ochthoecini appears in Fjeldså (2012) but there it does not fulfill requirements for new family group names in the ICZN (e.g. articles 13.1 and 16.1: no description or diagnosis, no explicit intent to establish it as a new
name etc.).
Scotomyias, Ohlson, Irestedt, Batalha Filho, Ericson & Fjeldså, new genus
Type species: Myiobius flavicans P. L. Sclater, currently recognized as Myiophobus flavicans
Included species: Scotomyias flavicans, Scotomyias inornatus (Carriker), Scotomyias phoenicomitra (Taczanowski & Berlepsch) and Scotomyias roraimae (Salvin & Godman). These species are all currently recognized as members of Myiophobus.
Diagnosis: Small (11–13.5 cm body length), compact chat-tyrants with subdued colours but proportions similar to those of close relatives Silvicultrix W. Lanyon and Ochthoeca. All species have an orange or yellow semi-concealed coronal stripe (males only), but lack the prominent bright eyebrow stripe typical of all Silvicultrix and Ochthoeca species. Upperparts dull olive green or brownish, underparts dull olive yellow to dull yellowish white, with smudgy olive flammulations on chest and body sides. Wings and tail have varying degrees of diffusely demarcated
cinnamon or buffy edges to secondary coverts and fringes of the remiges, forming distinctive wing-bars only in M. roraimae. These markings extend along the whole length of the feathers, with no contrasting black area at the base of the secondaries in the closed wing, as seen in Nephelomyias and Myiophobus. Iris always dark, legs dark grey
or blackish, upper mandible blackish, lower mandible usually dull orange to flesh-coloured. There are no known synapomorphies in internal anatomy for this group.
Cladistic definition: All descendants of the most recent common ancestor of Scotomyias flavicans and S. roraimae.
Habitat and distribution: All species inhabit understorey and thickets in humid forest in the Andes and the Guiana Highlands, mainly in the lower montane forest, up to 2700 m. Generally rather quiet and sluggish and normally found in pairs or family groups. Usually forage independently and do not follow mixed feeding parties; forage with short aerial sallies from a low perch.
Etymology: Gr. σκοτος skotos darkness, gloom; Mod. L. myias flycatcher. Refers to the dark and shadowy forest interior habitat of all species in the genus, a habitat in which few other members of Fluvicolinae are found. The
name is masculine in gender.
Comments: These species form a strongly supported clade sister to Silvicultrix (Fjeldså et al. 2018). In view of the close similarity both in plumage and syringeal characters (Lanyon 1986) and the fact that S. inornatus and S. flavicans replace each other geographically, we also infer that S. inornatus should be included in this genus, although genetic data is lacking for it.
Perso, I would have liked a generic separation of Fluvicola pica and Fluvicola albiventer, I don't know "Rheotriccus"
Empidonax could split into 4 genera if its polyphyly is strongly supported (Empidonax, Cnemonax and 2 unnamed genera)
Ugh...Empidonax are so similar to one another I would be strongly against any move to split up the genus. It's also a very useful term to use when talking about flycatchers. I would rather merge other genera in with the group as needed.
In the case of Empidonax, phylogenetic resolution and statistical support for non-monophyly is insufficient for taxonomic changes to be made at this point. There are no known diagnostic morphological features to distinguish
between the four Empidonax clades, but each of them possesses at least one unique ecological trait that separates them from their congeners (Johnson & Cicero 2002), as follows. Empidonax virescens (Vieillot, 1818) is the only species in its clade, and is also the type of the genus. It is unique in inhabiting bottomland forest interior, often near small streams. The second clade [E. flaviventris (W. M. Baird & S. F. Baird), E. flavescens Lawrence, E. difficilis (S. F. Baird) and E. occidentalis Nelson] differs from other Empidonax species by placing a mossy nest on a ledge or in a crevice, instead of placing a nest of plant fibers in a branch fork. The third clade, which consists of E. albigula is (P. L. Sclater & Salvin), E. alnorum (Brewster) and E. traillii (Audubon), differs from other Empidonax species by breeding in damp, often semi-open habitats like moist thickets and bog margins, in contrast to the generally dry to mesic woodland or forest edge habitat of other species. The fourth clade [Empidonax atriceps Salvin, E. fulvifrons (Giraud Jr.), E. minimus (W. M. Baird & S. F. Baird), E. wrightii S. F. Baird, E. hammondii (Xántus), E. affinis (Swainson) and E. oberholseri A. R. Phillips] differs from other Empidonax clades by having unmarked eggs. If a division of Empidonax into several genera is shown to be necessary, the name Empidonax is applicable to the clade containing E. virescens, whereas the name Cnemonax Brodkorb, 1936 (type species Empidonax atriceps) is available for a clade containing E. atriceps (clade 4 in this work). Any further new genera would need new names.
They apparently treated (Ochthoecini + Contopini + "Xolmini") as a trichotomy.I find it weird that they propose a linear classification of Fluvicolini - Ochthoecini - Contopini - Xolmini, where their clades are ( Fluvicolini + ( Contopini + ( Ochthoecini + Xolmini )))
G. Kohler's 2017 thesis (referred to in #75) includes the following new genera:
Inambariornis Cohn-Haft, Kohler, Aleixo, Brumfield & Ribas. Type: Hemitriccus spodiops (von Berlepsch, 1901).
Lanyonia Cohn-Haft, Kohler, Aleixo, Brumfield & Ribas. Type: Poecilostriccus senex (von Pelzeln, 1868).
Andinotriccus Kohler, Cohn-Haft, Aleixo, Brumfield & Ribas. Type: Hemitriccus granadensis (Hartlaub, 1843).
Bornscheinia Kohler, Cohn-Haft, Aleixo, Brumfield & Ribas. Type: Hemitriccus orbitatus (zu Wied, 1831).
Campina Cohn-Haft, Kohler, Aleixo, Brumfield & Ribas. Type: Hemitriccus inornatus (von Pelzeln, 1868).
Krotalotriccus Kohler, Cohn-Haft, Aleixo, Brumfield & Ribas. Type: Poecilotriccus capitalis (P. Sclater, 1857).
Physatriccus Kohler, Cohn-Haft, Aleixo, Brumfield & Ribas. Type: Poecilotriccus sylvia (Desmarest, 1806).
This work is (presumably) not published in the sense of the ICZN.Of course, a name introduce in a dissert is not available in accordance with the ICZN
This work is (presumably) not published in the sense of the ICZN.
Theses and dissertations (nowadays) are most often issued in a way that does not satisfy the criteria of publication set by the ICZN. But there is nothing in the Code that states specifically that names in theses or dissertations are not available. If a reasonable number of identical copies of this thesis were printed and made obtainable by the public, it would be published, and the names might in principle be deemed available.
(But the author of all of them would probably be Kohler alone.)
Has this since been published?The Pleistocene origin and diversification of the genera Colopteryx at 1.82
Myr (CI: 1.27-2.38), Lophotriccus and Oncostoma at 1.62 Myr (CI: 1.08-2.14) and
Inambariornis at 1.82 Myr (CI: 1.27-2.38) are discussed in detail on a separate
publication (Kohler et al. in. prep)