Alan
You determined for me to whittle away my evening?
You are in good company in failing to perceive this as a Tolmomyias, the authors of the first Alta Floresta inventory (Zimmer et al. 1997) missed the species due to then unfamiliarity with the vocalisations - see discussion in Lees et al. (2013). This despite the fact that it is a nuclear canopy flock species present in virtually every assemblage.
If my recordings don't convince you compare them with:
http://macaulaylibrary.org/audio/126754
http://macaulaylibrary.org/audio/127414
http://macaulaylibrary.org/audio/115022
http://www.xeno-canto.org/18058
http://www.xeno-canto.org/94647
We assigned the birds south of Alta Floresta to the nominate (with the disclaimer that we didn't know what calamae sounded like), the soils are a mosaic and it is likely that both are in contact somewhere between the two nearest localities (c. 50km) over which there is no geographic barrier (Lees et al. 2008). So yes, there is an excellent chance they could be sympatric and likely to be behaving as BSc species given that their song types are well preserved on either side of this zone.
"paraensis"
You determined for me to whittle away my evening?
The first cut of "paraensis" is certainly very unusual for a Tolmomyias - I was reminded of something and have just remembered the "song" of Todirostrum pictum, which is not dissimilar (though the notes are delivered at a faster pace). The second song is also unusual and sounds a little like a Myiarchus!
You are in good company in failing to perceive this as a Tolmomyias, the authors of the first Alta Floresta inventory (Zimmer et al. 1997) missed the species due to then unfamiliarity with the vocalisations - see discussion in Lees et al. (2013). This despite the fact that it is a nuclear canopy flock species present in virtually every assemblage.
If my recordings don't convince you compare them with:
http://macaulaylibrary.org/audio/126754
http://macaulaylibrary.org/audio/127414
http://macaulaylibrary.org/audio/115022
http://www.xeno-canto.org/18058
http://www.xeno-canto.org/94647
I think in the Cotinga article, it is suggested that the nominate form (or a bird with identical song) is also present, so assume this is the second taxa. I would guess the edaphic gradient is not a simple line so might these birds be locally sympatric anyway? I think the same happens with a large non-passerine, with two song types on different soils, that someone is supposed to be working on..
We assigned the birds south of Alta Floresta to the nominate (with the disclaimer that we didn't know what calamae sounded like), the soils are a mosaic and it is likely that both are in contact somewhere between the two nearest localities (c. 50km) over which there is no geographic barrier (Lees et al. 2008). So yes, there is an excellent chance they could be sympatric and likely to be behaving as BSc species given that their song types are well preserved on either side of this zone.
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